PERIODIC REORGANIZATION OF P. CAUDATUM 



73 



we have described for Paramaecium aurelia, i.e., animals with 

 only one micronucleus and a completely destroyed macro- 

 nucleus, the remnants of which are scattered in the cell. This 

 one micronucleus, as in Paramaecium aurelia, is the carrier of 

 the life of the race through the new rhythm. 



Maupas and Calkins in the conjugation of Paramaecium cau- 

 datum figure eight micronuclei, the products of the syncaryon, 

 which are formed immediately after the separation of the con- 

 jugants. This we have verified in conjugation (fig. 25). In the 

 reorganization process we have discovered animals at a slightly 

 later stage with four micronuclei and four macronuclear anlagen, 

 the latter representing four transformed micronuclei. The 

 micronucleus which persists and carries the life of the race there- 



• 



d.h.l 



a.h.2, 



d.h.3. 



ni.H. 



Text fig. 5 



fore must have undergone three divisions producing the four 

 micronuclei and four macronuclear anlagen in the stage under 

 discussion. The micronuclei (fig. 19) are characterized b}' 

 their granular structure and somewhat glistening appearance. 

 In the preparations the few granules appear pure blue without 

 any reddish tinge. These micronuclei resemble, as far as one 

 may judge from figures, one of the four micronuclei which Klitzke 

 ('14, text fig. C, copied in the present paper as text fig. 5) 

 shows in the conjugation of Paramaecium caudatum. This 

 micronucleus (Klitzke's 'Micronucleusanlage,' d.k. 3) he believes 

 undergoes degeneration together with two other micronuclei 

 (d.k. 1, d.k. 2). The marked differences he points out are not 

 observable in our specimen in the reorganization process, but 

 this may be due to the fact that the animal has not yet reached 

 this point. Or it may be that no micronuclear degeneration 



