554 Theophilus s. Painter, 



period. In the first maturation division he describes thirteen bivalent 

 chromosomes, one of which must be the accessory though he was 

 not able to identify this body. II. While he states that he could 

 not distinguish the accessory body from the other chi-omosomes, he 

 adds, "We can say positively that the whole bivalent hétérochromosome 

 does not pass undivided into one of the second spermatocytes" 

 (p. 180), III. He describes, in the spermatogonia! divisions, the 

 presence of two very small chromosomes, the history of which he 

 could not follow with certainty. 



"Two of these spermatogonic chromosomes are very small (S); 

 the subsequent history of these could not be ascertained with any 

 degree of certainty" (p. 175). Again, "In the equatorial plate of the 

 first spermatocyte (figs. 53, 54) are found thirteen larger bivalent 

 chromosomes, and sometimes a minute chromatin body (S); which 

 does not appear to be bivalent, at least it is not bipartite. The 

 latter may represent one of the two minute chromosomes of the 

 sperraatogonial (fig. 41); one of these small bodies is occasionally 

 found in the monaster stage of the second spermatocyte (figs. 62, 63)" 

 (p. 178). IV. He states, "There is no rest stage in the spermato- 

 cytic history" (p. 175). 



Wallace found, in Agalena 7taevia, that the accessory chromo- 

 some arose from two spermatogonial chromosomes, and in this respect 

 her work agrees with that of Montgomery, but she found that the 

 accessory elements passed to one pole in the first maturation mitosis, 

 a behavior which had been described by Berry in an earlier work. 

 It seemed altogether probable, therefore, that essentially the same 

 behavior would be found for the Lycosidae when they were re- 

 examined. From Montgomery's description, it seems probable that 

 he had in Lycosa insopita either two microchromosomes or else two 

 planosomes. 



Spermatogonial division stages that are suitable for study 

 are lacking from the material examined so that it has not been 

 possible to follow the behavior of the accessory elements during this 

 period. 



Contrary to Montgomery's description for Lycosa insopita, there 

 is a very distinct rest period for Lycosa communis (Fig. 114). In 

 this figure we see, besides the large nucleolus, which represents 

 the accessory elements, two very small nucleoli (F). These persist 

 as distinct deeply-staining bodies during the whole growth period. 



In the first division, the accessory cliromosome lies on the outside 



