REPRODUCTIVE SYSTEM OF FREE-MARTINS 475 



the fold of peritoneum which overhangs the anterior part of the 

 gonad (fig. 14). This position of the Wolffian duct is character- 

 istic of the male. As for the Miillerian ducts, the tube is en- 

 tirely atrophied. The horns of the uterus are present and like 

 those of a normal female, in histological structure, but much 

 smaller in diameter. However, in the preparation studied, they 

 do not unite, although they were sectioned posterior to the 

 point at which they normally unite in the female. 



In the 22.5 cm. free-martin T4, the epididymis and anterior 

 part of the Wolffian duct resemble those structures in free-martin 

 2. The convoluted portion of the Wolffian duct (figs. 15 and 16). 

 lies in the fold of peritoneum overhanging the gonad. Poste- 

 rior to the gonad, the Wolffian duct ends in an enlargement. The 

 caudal part of the Wolffian duct is present and, as in case 13, the 

 lumen is discontinuous. The entire tubular portion of the Miil- 

 lerian duct is atrophied. In the location where the anterior part 

 of the horns of the uterus are found, there is a large structure 

 which may be a rudiment of one of them. It has a typical, epi- 

 thelial wall and is about the diameter of the normal uterine 

 horn. It is closed at both ends, having no connection with any 

 other structures. Lateral to the posterior ends of the Wolffian 

 ducts are coiled tubules, one on each side. These are presum- 

 ably the anlage of seminal vesicles which develop in the male 

 from the Miillerian ducts at their posterior end. 



Summary. The gonads of the free-martins measuring 20 cm., 

 21.5 cm. and 22.5 cm. resemble the testis in the entire absence of 

 cortex, the appearance of the albuginea, peritoneum and rete, 

 and the connection of the rete with the Wolffian duct through the 

 Wolffian tubules. They resemble the ovary in the position of the 

 rete and in the structure of the medullary cords, with the excep- 

 tion of the few sex cords in T2 which have the appearance of 

 seminiferous tubules. T2 resembles the male in the condition of 

 both ducts. T13 and T4 resemble the female in the partial 

 atrophy of the Wolffian duct and resemble the male in the 

 atrophy of the oviduct, and non-union of uterine horns. 



