685 
of the Nebenkern as in the Salamander’s spermatocytes but lying free in 
the cell body, and even at the early stage represented in the figure already 
showing signs of being individually duplicated (fig. 1). 
Except in the incorporation of the small chromatic body (fig. 1c) 
into one or other of the chromosomes, the ensuing division is a perfectly 
normal heterotypical one, and, as I have said, the primary nucleus as well 
as the daughter elements (spermatids) each contain 8 chromosomes. So 
that although there is apparently a reduction in the number of the 
chromatic elements as compared with the previous division, the reduction 
would appear more comparable to the type described by GUINGNARD 
during the formation of vegetable pollen, than to that pointed out 
by Hertwie in the spermatocytes of Ascaris. 
Both the Nebenkern (Archoplasm) and the Lesser Nebenkern are 
reabsorbed during the division. But while a new archoplasm is itself 
regenerated in the daughter elements (spermatids), as a coalescence of 
spindle fibres, the Lesser Nebenkern does not again appear. 
It should moreover be noted that the spindlefibres of each division, 
are not formed out of the existing Nebenkern (Archoplasm), which is 
Fig. 1. Fig. 2. 
absorbed into the kytoplasm, but that new archoplasms are formed 
from successive crops of spindle fibres. 
When first formed the spermatids are spherical, and their nucleus, 
under the action of reagents, is intensely chromatic. In their kytoplasm 
there are numbers of large granules, presumably equal to those de- 
