43 
3) The kidney of fishes consists of two quite distinct parts 
(1) the secreting tubules and Malpighian bodies, and (2) an adenoid 
intertubular material. 
4) The head-kidney in adult Teleosts is not a “specialised” 
portion of the primitive kidney, but simply an accumulation at the 
anterior end of the organ of the lymphatic tissue which exists between 
the tubules throughout the kidney. 
5) There is, so far as I can make out, no anato- 
mical or physiological relationship of any kind be- 
tween the suprarenals in Teleosts and the lymphatic 
head-kidney. 
Again, in the “Anatomischer Anzeiger” Band XII, No. 9 and 10, 
1896, in a paper “On the so-called Suprarenal Bodies in Cyclostomata” 
written in conjunction with Mr. WALTER E. COLLINGE, dated July 30th 
1896, I reiterated my opinion that there is no connection between 
suprarenals and head-kidneys. We pointed out that “just as RATHKE’s 
theory as to the homology of the modified head-kidney and suprarenal 
bodies crept into a number of the earlier text-books, so, brought for- 
ward again by WELDon, it has had a like career”. 
Our conclusions as far as they bear upon the matter in hand 
were as follows: — 
“As yet there is no evidence which would lead us to 
suppose thatthe pronephros of Cyclostomata represents 
the suprarenal bodies of Gnathostomata, and the theory 
put forward by RATHkE, and reiterated by WELDON, has no 
evidence in its favour. We would emphasise the fact, 
that evidence is still wanting to prove that there is any 
relation, beyond that of position, between the supra- 
renal bodies of Vertebrates and the renal organs, the 
two having widely different functions, which in no way 
ever become modified so as to replace one another. 
Up to the present we are inclined to regard the 
suprarenals as paired bodies first appearing in Fishes 
and increasing in importance as we ascend the scale, 
and not in any sense as the remains of a structure with 
a past history. One or other, of the constituent parts 
is present if not both in all Vertebrates with the pos- 
sible exceptions of Cyclostomata and Dipnoi.” 
