428 
gives rise to the usual ciliary nerve. It is to be noted that the 
sympathetic commissure X to V + Vllisexternal to the membranous 
ear and that in this position Pınkus [6] found in Protopterus a 
bundle of lateral line fibers passing from VII to X and IX and 
that Strong in the tadpole found a bundle of general cutaneous fibers 
from IX to VII. 
All of the fibers of the V—VII except the sympathetic leave the 
ganglionic complex in five groups composed as follows: 
1. Ramus hyomandibularis. (a) Motor VII. (b) Fasciculus communis 
fibers from the geniculate ganglion distributed to naked end buds, 
lying for the most part along the course of the opercular canal, but 
external to it. (c) Lateralis fibers, comprising the whole of the 
ventral one of the two acustico-lateral roots and innervating the canal 
organs of the operculo-mandibular canal. (d) General cutaneous fibers, 
curving backward from the Gasserian ganglion. These in their peripheral 
distribution correspond to the fibers which STRONG finds emerging 
with IX in the tadpole and forming the commissure IX to VII. 
2. Ramus palatinus. Fasciculus communis fibers from the geni- 
culate ganglion, distributed exclusively to taste buds of the mouth. 
3. Ramus recurrens VII. Commonly called ramus recurrens V, 
but, as the fibers are all fasciculus communis, that name is inappro- 
priate. These fibers arise from the geniculate ganglion in several 
strands, pass dorsad in the meninges and finally emerge on the dorsal 
surface of the head and then turn caudad into the trunk. They 
anastomose with each other and farther back with the dorsal ramus of 
each spinal nerve. I have not been; able to demonstrate that they 
supply either canal organs or end buds. They have not, however, 
been traced caudad more than a few segments into the trunk. They 
clearly correspond to the superficial, or accessory “lateral line” nerves 
of the Gadoids. 
4. Ramus ophthalmicus superficialis. This includes the r. oph. 
sup. VII, arising from the dorsal one of the two lateral line roots 
and distributed to the supra-orbital canal, and the r. oph. sup. V, 
from the Gasserian ganglion for the skin above the eyes. The latter 
communicates in front of the eye with the general cutaneous fibers 
of the r. maxillaris V. The r. oph. sup. VII and the r. oph. sup. V 
are fused throughout their entire course, but each can be easily 
distinguished and separately followed by reason of the difference in 
the calibre of the fibers. 
The r. oticus follows the r. oph. sup. for a short distance. 
When it separates it draws off both lateral line and general cutaneous 
