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fibers. The former innervate the single canal organ of the main canal 
of the head between the operculo-mandibular canal and the infra- 
orbital and the first two organs of the infra-orbital line. Between 
the latter organs, however, the infra-orbital canal opens out, so that 
the second organ is a naked papilla. The general cutaneous fibers 
anastomose with others from the r. oph. sup. V and supply the skin 
adjacent to the three canal organs just mentioned. 
5. All of the other fibers of the V—VII complex fuse into a 
common infra-orbital trunk which can, however, be analyzed into its 
elements both proximally and distally. (a) R. buccalis, from the 
dorsal lateral line root and distributed to the infra-orbital canal. This 
canal opens out and disappears in the ventral part of its course; 
but, like the lateral line of the trunk, is represented by a row of naked 
buds which receive their innervation from the coarse fibered r. buccalis. 
(b) General cutaneous fibers from the Gasserian ganglion. These 
represent the r. maxillaris V and the r. mandibularis V. 
(c) Fasciculus communis fibers from the geniculate ganglion are clearly 
seen to enter this trunk. Certain fibers from the r. maxillaris supply 
the lining of the mouth and are probably derived from this com- 
ponent (chorda tympani?). (d) The motor V. 
Broadly, then, we find the fasciculus communis component to 
conform to the descriptions of the other recent writers who have 
treated of it in the Ichthyopsida — Srrone [8] in the tadpole, 
ALLIS [1] in Amia, Kınassury in Necturus [4] and fishes [5], 
though it should be noted that a “lobus trigemini” such as is described 
by Kryessury for Amiurus [5] is totally wanting in Menidia. The 
same agreement prevails respecting the general cutaneous component. 
Regarding the acustico-lateral system, there is not the 
same unanimity. I confirm the conclusion .reached by Potnarp [7] 
in the Siluroids that the lateral line system is to be regarded as a 
single organ without direct affinities with any of the nerves which it 
may accompany in its exit, save that I would include with it the 
VIII nerve. In this again I agree with the other writers just mentioned 
and with Core [2]. though I dissent from the latter’s association of 
the lateral line system with the VII nerve. The reasons which he 
gives for excluding it from the vagus apply equally mutatis mutandis 
to the facial. 
COLLINGE in his most recent paper on the Teleosts [3] comes to 
a totally different conclusion, as indicated in his concluding paragraph: 
— “It will be seen from the above tabular view of the nerves that 
the innervation [of the canal organs| proceeds very largely from the 
