THE GERMAN CARP IIST THE UNITED STATES. 533 



It seems probable, however, that the character of the scales should 

 be placed with the other four categories of characters given above 

 as being another moditication brought about by artificial breeding and 

 selection and not as a condition due simpi}' to conditions of domesti- 

 cation, as is sometimes supposed. All of these characters are probably 

 heritable, although some of them, such as rate of growth and time of 

 sexual maturity', may undoubtedl}' be readily inliuenced by external 

 conditions in the individuals of a single generation. Furthermore, 

 there apparently can be all combinations of these characters, and the 

 so-called different varieties and races are the fish possessing the vari- 

 ous combinations. In general, it maj^ be said that the most highly 

 specialized carp are those which are destitute of scales, which grow 

 quickly, are high in proportion to their length, and tend to have a hump 

 back of the head, and which become sexually mature at an early age. 



These various forms of carp probabl}- difl'er in no essential way, 

 except that they are not so well ditferentiated and established, from 

 what are spoken of as "breeds" by stock breeders. There would 

 appear to be no valid reason for calling those with the different char- 

 acter of scales "varieties," and to class those which are differentiated 

 as to form as "races." It is merely that the most obvious characters 

 are those which have become most permanent Ij^ established hj selec- 

 tion, namely, character ol scales first and form second. Walter 

 claims that ability for cpiick growth has also been fixed in certain 

 stocks. Thus a fish of good quick-growing stock may later make a 

 good growth even if poorly nourished during its first or second 3'ear, 

 whereas a fish of poor stock under similar conditions would be perma- 

 nently stunted. The hardiness, or ability to resist climatic condi- 

 tions, he snys has not j^et been made permanent in any stock, though 

 it is claimed that scale carp possess the abilit}^ to a greater degree than 

 the others. The adaptability to climatic conditions probably becomes 

 reduced rather than increased as the other characters are developed. 



All intermediate stages are found in the sets of characters men- 

 tioned. For example, fish may be entirely covered with scales, but 

 the scales are larger and fewer in number than on the regular scale 

 carp, and, similarly, one finds all gradations between the leather and 

 the mirror carp. The same thing is true of the form of the body. 

 This is especial!}' the case with the fish in our waters, where all kinds 

 have become established and have interbred until there is a complete 

 series in the gradation of characters in almost an}^ lot of fish taken, 

 and a division of them into varieties must be an arbitrary one. As a 

 matter of convenience in my work, those fish which had larger and 

 fewer scales than typical scale carp I called mirror carp. Some 

 authors state that the leather carp should be entirely destitute of scales; 

 others that it may have a row of scales along the back and a row on 

 each side. In no case in the Great Lakes did I see a carp entirely 



