445 



Fig. 4. 

 c. t. n, eos. l. c. t.f. c. t. n. 



/ 



:,*T »■ 





5«'^..;.-,/ 



' \ ' ~ - eos. l. 



eos. l. c, t.f. c. t.f. 



Fig. 4. Stratum compactum from gastric mucous membrane (estuary salmon); to 

 show its continuity with the neighbouring connective-tissue fibres, and the number of 

 eosinophile leucocytes about it. (The details of the nuclei of the leucocytes have been 

 omitted.) Hematoxylin and eosin. 



Oil the projections between the mouths of the glands, they have a nar- 

 row dear hem on the surface, and below that a portion of the cell, 

 which looks more hyaline and stains rather more deeply than the rest, 

 and is sharply marked off from the deeper portion, which shows an 

 ordinary protoplasmic structure. These cells are not goblet-cells, and 

 no such cells are found in the stomach. The glands are very thickly 

 set over the stomach and open by wide ducts, down which the super- 

 ficial epithelium extends unaltered for some distance, usually down to 

 the point where the glands begin to branch. Oppel notes that this 

 branching or opening of several secreting tubes into a common duct 

 occurs very frequently in the trout, and I think that in the salmon 

 it is still more frequent. 



2) Before the branching of the glands begins, the characteristic 

 upper end (Oberende, Oppel) of the cell becomes less distinct and 

 soon ceases to be evident, and the whole of the cell body becomes 

 granular and undifferentiated. This I call the intermediate epithelium 

 (Fig. 3). In the glands of the cardiac or anterior part of the stomach 

 this part of the gland is very short, but in the pyloric region it occu- 

 pies almost the whole of the gland. 



3) The zymin-secreting epithelium cells do not exhibit any trans- 

 ition from the intermediate ones, but are sharply distinguished from 

 them (Fig. 3). They are cubical, have a large rounded nucleus poor 



