78 BLAKE. NEOMILLSPAUGHIA 
precisely with Coccoloba, and the differences indicated are so 
comparatively slight and so weakened by intergradient forms 
that they can not be considered of generic value. 
Of the five species of Campderia described up to 1890, repre- 
senting three valid species, all are accounted for by Lindau in his 
monograph of Coccoloba. A plant of very different characters, rep- 
resenting an undescribed genus, has more recently been described 
from Honduras by Donnell Smith* under the name Campderia 
paniculata, and with it is clearly to be associated another species 
from Yucatan described by Grossf in 1913 as Podopterus emar- 
ginatus. Both of these are shrubs or trees with orbicular leaves, 
cordate at base and strongly emarginate at apex. The inflores- 
cence is a terminal panicle compounded of numerous slender 
racemes, the branches minutely scaly-bracted at base. Such an 
inflorescence is found in only two species of Coccoloba, forming the 
section Paniculatae of Meisner. It is in the perianth, however, 
that the chief distinguishing characters of this new genus appear. 
The tube is very short, and the five segments are in two distinct 
series; the three outer are distinctly winged from apex to base, 
and the wings are slightly decurrent on the pedicels. The two 
inner are flat, wingless, and shorter than the outer. In fruit all 
are somewhat accrescent, dry, and entirely free from the achene. 
In Coccoloba, on the other hand, the five perianth-segments are 
similar or subsimilar, more or less fleshy or coriaceous, in fruit 
flat or rarely slightly carinate but never winged, and usually 
adherent to the achene. The affinity of the new genus is clearly 
with Podopterus rather than with Coccoloba. From the latter it 
- may be distinguished by its paniculate inflorescence, more nat- 
rowly winged perianth, and slightly winged scarcely elongated 
fruiting pedicels. In Podopterus the flowers are borne in dense 
fascicles at the tips of very short branchlets or in short axillary 
racemes, and the perianth and the fruiting pedicels are very 
strongly winged. The exact nature of the inflorescence in the 
type species of Podopterus, P. mexicanus, not clearly described by 
previous authors, deserves a word of explanation. The leaves 
are alternate and —— bear fascicles of two to four similar ones 
_* Bot. Gaz. 27: 440. 
+ Rep. Nov. Sp. Hae se ms 1913. 
