SPERMATOGENESIS IN AMPHIBIA AND INSECTS 3 
Terni (14). The idiosome and Golgi apparatus have been often 
described by earlier workers. Together they form much the 
most conspicuous object in the spermatocytes. The idiosome 
appears as a more or less irregularly spherical mass, staining 
rather more darkly than the general cytoplasm. ‘To its surface 
are applied a large number of small, separate Golgi rodlets, each 
shaped somewhat like a banana (fig. 1), and seemingly never 
interconnected to form a ‘network.’ These rodlets, now defi- 
nitely shown to be Golgi elements, have of course been long known 
under a variety of names, e.g., the ‘Archoplasmaschleifen’ of 
Hermann, the ‘Centralkapsel’ and ‘Pseudochromosomen’ of 
Heidenhain, and the ‘formazioni periidiozomiche’ of Terni. 
During the maturation divisions the idiosome goes to pieces 
and its exact behavior during the division period is unknown. 
The Golgi rodlets undergo a great loss in staining capacity, and 
appear to be simultaneously fragmented, so that with the usual 
methods they cannot be satisfactorily followed in the division 
stages. It is certain, however, that the Golgi pieces (dictyo- 
somes) are collected about the poles of the spindle (fig. 2) as 
described in invertebrates by several workers, although the exact 
method by which this distribution is achieved has not yet been 
clearly made out. At the close of the first maturation division 
the idiosome is reconstituted with the Golgi rodlets applied to its 
surface as before (fig. 3). Similarly, in the spermatids the same 
structure is built up soon after the second maturation division 
is completed. In my preparations, however, the Golgi rodlets, 
as distinct elements, are soon lost, and the outer part of the 
acroblast (as the Golgi apparatus plus idiosome may now be 
called) tends to impregnate rather uniformly with the various 
Golgi methods (figs. 5 and 6). 
The acroblast lies at first some little distance from the nucleus; 
but presently it moves over into contact with the nuclear mem- 
brane and a small clear vesicle makes its appearance on one side 
of the acroblast (fig. 5). This is the “Sphaerenblaeschen’ of Meves 
and the acrosome of McGregor. In general appearance it closely 
resembles the homologous structure which I have described in 
the hemipteran spermatid (Bowen, ’20), and for which I have 
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