INEQUALITIES IN HOMOLOGOUS CHROMOSOMES 125 
to one colored, while some give all colored again. This shows 
that there are two factors concerned. The presence of both is 
needed to produce the colored flowers, the absence of either one 
giving white. It is the F, result of an ordinary dihybrid Men- 
delian ratio, in which there are nine cases where both factors are 
present and colored results, three cases where only one factor is 
present and white results, three cases where the other factor is pres- 
ent and white results, and one case where neither factor is present 
and a white which breeds true results. 
The colored individuals in the F, generation are found to be- 
long to six classes depending upon the presence or absence of a 
purple factor, a light-wing factor giving a bi-colored flower, and 
a factor for intensity of pigmentation, in the absence of which a 
dilutely colored flower results. The wild type is intensely colored, 
and bi-colored,i.e., purple with blue wings. By dropping the pur- 
ple a red bi-colored or uniformly colored flower is obtained in both 
the dilute and intense series, by dropping the bi-color factor uni- 
formly purple or red are obtained, by dropping the intensity 
factor both varieties of the red are obtained in dilute form, and, 
finally, by dropping either the color-producing base or the color 
developer we get any of these color varieties in the albino form. 
The wild purple Sicilian species contains all these factors. By 
the dropping of these factors one by one and by inbreeding, 
‘all the color varieties of our domestic sweet peas have been 
obtained. 
Now, it seems to me that it would be quite possible to account 
for phenomena of this kind in plants and animals as the result 
of an unequal division of the chromosomes in the reduction 
division, similar to what I have found evidence of in Tettigidea 
parvipennis. If but one member of the pair of chromosomes 
showed the deficiency it might not give a result in the organism. 
When both chromosomes of the pair show such a deficiency, a 
condition which would result only when inbreeding occurs, then 
such a deficiency might be shown in the somaplasm by some such 
defect as albinism, blackness, dwarfishness, ete. It is true that 
such a chromosome would evidently be an abnormality of a 
deficient kind, but are not all such traits as albinism in plants 
