320 SCOTT. [Vol. XI. 



The formnina of the skull are antelopine rather than cervine 

 in character, which is doubtless due to the backward shifting 

 of the orbit, this displacement having proceeded farther than 

 in the deer, especially the hornless genera. The infraorbital 

 foramen is single and occupies rather a posterior position ; it 

 lies above the interval between A? and p_4 (in the deer it is 

 above or in advance of P_£) and is roofed by the tubercle in 

 which the masseteric ridge terminates. The optic foramen is 

 lower than in most antelopes, in correspondence with the less 

 elevated position of the orbit. The foramen lacerum anterius 

 is rather small and rounded, while in the deer it is very large 

 and of irregular shape, especially so in MoscJuls, where it forms 

 a great fissure. As in the Pecora generally {and indeed most 

 artiodactyls) the foramen rotundum is not present, though its 

 position is in some specimens marked by a pair of minute 

 openings. The foramen ovale is quite large and placed in- 

 ternally to the glenoid cavity ; it varies in shape, being in some 

 individuals nearly round and in others narrow and elongate. 

 In spite of the very small size of the auditory bullae, the fora- 

 men lacerum medium and posterius are mere slits, much nar- 

 rower than in Cariaais, and the bulla is not channelled by the 

 carotid canal. A postglenoid foramen is present, though of 

 small size. The condylar foramen occupies a position where it 

 is concealed by the prominence of the inferior portion of the 

 condyle. The squamosal is perforated by two large vascular 

 openings above the root of the zygomatic process. The re- 

 markable position of the posterior palatine foramina has already 

 been described ; it only remains to mention an irregularity in 

 the two sides which sometimes occurs, — thus, in a male skull 

 the left foramen is fully a quarter of an inch behind the right. 



The second type of skull is that of the male, the knowledge 

 of which we owe to Osborn and Wortman. Their beautifully 

 preserved specimen is still by far the best that has been dis- 

 covered, and is one of the most remarkable and curious of 

 mammalian fossils. The striking resemblance of this skull to 

 a miniature Uintatheriimi has been commented on by the 

 authors mentioned, and is an excellent example of a superficial 

 parallelism between two forms which are as widely separated 



