No. 2.] ENTEROxW OF AMERICAN GANOIDS. 42 I 



Ciliated epithelial cells have been found in the oesophagus of 

 sturgeons, but no one mentions their presence in the stomach, 

 and several deny their ever existing there. 



Concerning the glands in the stomach of the sturgeon Ley- 

 dig says : " They are short cylindrical sacs . . . lined with 

 great regularity by a clear and delicate cylindrical epithelium, 

 which is continuous at the edge of the gland orifice with that 

 of the surface epithelium; the cells of the two cylindrical epi- 

 theliums are distinguished in this, that in the surface epithe- 

 lium they are larger and distended towards their free extremity 

 by a molecular mass." If the above statement applies to all 

 the glands of the stomach, Leydig's specimen differed greatly 

 from the one studied by me. It has already been noted that 

 in my specimen the greater part of the gastric epithelium is 

 ciliated. Among the ciliated cells are numerous beaker-cells, 

 many of which are open at the free end. The glands of the 

 stomach are of the two ordinary kinds, cardiac and pyloric; 

 the latter occupy but a small area compared to the former. 

 The cardiac glands are differentiated into two very clearly 

 recognizable portions, a superficial or mouth part and a deeper 

 or body portion. The relative length of the two parts through 

 the middle portion of the stomach is as i to 3 nearly (Fig. 9). 



On both sides of this area the glandular cells are gradually 

 replaced by those of a more nearly cylindrical form, till finally 

 the tubules are lined by the latter form of cells only. In all 

 the tubules, so far as observed, except those in the pyloric 

 region, the mouth portion is lined by ciliated columnar cells 

 (Figs. 6, 8, 9). No difference was detected between these and 

 the cells of the surface epithelium except that the latter are 

 longer and their attached ends do not end so abruptly. Beaker- 

 cells are found in the mouths of the glands as well as on the 

 free surface. The nuclei of the epithelial cells stain deeply 

 with haematoxylin ; the cell-body stains sparingly with eosin 

 unless the stain is left on for some time; this applies equally 

 well to the gland cells proper. The part of the cell next to 

 the so-called basement membrane stains more deeply than the 

 part next to the lumen of the gland. All the cells in the 

 glandular portion of the tubule have approximately the same 



