No. 2.] ENTERON OF AMERICAN GANOIDS. 429 



ends can be traced down among the connective-tissue corpus- 

 cles of the mucosa; not the slightest indication of a basement 

 membrane was noticed. From many of the beaker-cells of the 

 oesophagus can be seen conical or rounded projections of 

 mucus, but, unlike Scaphirhynchops, these mucous masses are 

 stained very little. The transition from oesophagus to stomach 

 is gradual. Towards the gastric end of the oesophagus the 

 epithelium gradually infolds, forming short, broad follicles. 

 These are soon replaced by the true gastric glands of the stom- 

 ach. The latter are lined by large granular, irregularly cubical- 

 shaped cells, except the part near the exit of the gland upon 

 the surface, where the cells are of a broad cylindrical shape, 

 but there is no sharp line of demarcation between the two. 

 The glands of Lepidosteus, unlike those of the other forms, 

 have, for the most part, no portion lined by cells of the same 

 size and appearance as those of the surface epithelium. The 

 part corresponding in position to the mouth is lined by cells 

 two or three times as large as those of the surface. The rela- 

 tions of the cells at this point are such as to give the appear- 

 ance of the gland having been forced up through the surface 

 epithelium like a wedge (Figs. 23 and 26). 



As a rule the glands open singly upon the surface, but some- 

 times two or more have a common outlet. The cells along the 

 upper half of the glands show clearly the thread-like continua- 

 tions extending from their basal end ; in the deeper part of the 

 gland they are somewhat shorter. The nuclei of the gland 

 cells are circular, those of the surface epithelium oval in out- 

 line. No indications of cell division were noted. 



As we approach the pylorus the glands grow shorter; at the 

 same time the upper part becomes lined with cells like those 

 of the surface epithelium, till finally the glands are lined with 

 columnar cells only; these glands are considerably shorter 

 than the cardiac glands. In the cardiac region the glands are 

 close together, and of about the same diameter at all parts 

 (Fig. 26). The lumen is distinct and enlarged somewhat just 

 before it opens upon the surface. Cilia were not found in any 

 part of the lumen of the gland in any portion of the stomach. 



