384 



WHITMAN. 



[Vol. IV. 



nucleated ends next to the muscular wall. The reduction of the 

 lumen in this part is due to the development of this glandular 

 epithelium. It is in this glandular portion that the granular cor- 

 puscles {gc) which plug the spermatophore are produced. 



I have a series of sections of this region, showing the gland- 

 cells fixed in the very act of secreting these corpuscles. The 

 corpuscles are somewhat pyriform in shape, with the smaller 

 end tapering to a fine thread, which is connected with the cen- 

 tral end of the producing cell (Fig. 2, gc). As soon as the cor- 

 puscle is fully liberated, it assumes a more or less elliptical form. 

 These corpuscles sometimes nearly fill the whole lumen of the 

 duct ; sometimes they lie in masses that resemble clusters of 



Fig. 2. — Section from the glandular anterior half of the duct, in which the lining 

 epithelium (f/) is transformed into radial, pyramidal gland-cells, with nuclei at the 

 external bases. Granular corpuscles (^c) in process of formation. Muscular layer 

 thinner than in Fig. i. The lumen of the canal is very much reduced by the thick- 

 ening of the epithelial lining. 



blood-corpuscles. I find a few such clusters in the posterior 

 non-glandular region id), but I doubt if they move backward to 

 this part as a regular thing. They were more probably thrown 

 backward from the place of origin by irregular, peristaltic con- 

 tractions during the process of killing. At the anterior end (w) 

 of the corpuscle-secreting region, the glandular lining becomes 

 thicker, and passes imperceptibly into the still thicker lining of 

 the terminal horns. I find no corpuscles at this level (w), and 

 it is here that the free end of the spermatophore is evidently 

 secreted. The saccular part of the spermatophore, as before 

 noticed, is secreted in the horns (j-), while the narrower base is 



