R. R. Bensley 123 
epithelial cells frequently show a peculiar radial striation in the muci- 
genous border due to the optical projection into this border of struc- 
tures which are really on the surface of the cell. Carlier, 99, has shown 
that in the gastric epithelium of Triton the cement substance is not 
confined to the contiguous margins of the cells but extends in radiating 
lines over their free surfaces. This may be particularly well seen in 
sections of the newt’s stomach stained by the iron hematoxylin method. 
Similar features are also exhibited by the cement substance of the gas- 
tric epithelium of mammals. In longitudinal sections of the cells in 
iron hematoxylin preparations the cement lines are seen as black dots 
between the free edges of the cell, or as a line stretching across the 
cell near its free end, according to the focus. In the latter case fine 
lines of cement may be traced from the intercellular line in the direc- 
tion of the end of the cell over the convex surface of the mucigeneous 
border. At the point where these vertical 
lines join the cement line, there is a granule- 
like thickening, which might readily be mis- 
taken for the basal granule of a striated 
cuticle. The same appearances may be made 
out by careful focussing in a surface view of 
the ends of the cells where it is seen that the Fig. 5. Surface view of 
[ities OL cement over the surtace: of ithe cell) > Celsiof the suxtace opithe 
lium of the human stomach. 
> ; als rT WW ‘ ial < 1500. Stained in iron haem-— 
form an irregular network with nodal thick- 25 into Show Conont 
euines (Hig..5)./ This is undoubtedly the-ex:) nes Between the cells ana 
penawoen, ot the appearance represented, by. sauce which are contmucd 
Schaffer in his Fig. +4, where a row of gran- Cell. 
ules is shown along the free border. 
From the foregoing facts it is obvious that the differences between 
the fundus glands and cardiac glands are not fundamental. This‘is in- 
dicated by the occurrence in the cardiac glands of all‘the characteristic 
glandular cells of the fundus glands. Only the relative frequency of 
the various cells is altered. The cells which are most numerous are 
muciparous cells morphologically equivalent to the chief cells of the 
neck of the fundus gland and to the pyloric gland cells, although phy- 
siologically much less active than these. In view of the fact that the 
highly branched glands may be to a large extent formed of zymogenic 
chief cells which are the morphological equivalents of the chief cells of 
the body of the fundus gland but little importance can be attached to 
the mode of branching as a specific character of the cardiac glands. 
It now remains to be seen to what extent these conclusions can be 
supported by data derived from the examination of other mammalian 
stomachs. 
Cole, ae 
