288 EDWARD PHELPS ALLIS, JR. 
A ventral myodomic cavity is represented in those parts of 
the canales parabasales posterior to the foramina carotica. 
The antipterygoid is said by Gaupp (’00, pp. 541 and 542) to 
be the homologue of the ascending process of the quadrate of 
the Amphibia and to be wholly wanting in the cranium of mam- 
mals. The ala temporalis of the mammalian cranium is con- 
sidered by him to be represented, in reptiles, by the processus 
basipterygoideus. Fuchs (712, pp. 91 to 95), on the contrary, 
maintains that the antipterygoid (epipterygoid, Fuchs) is the 
homologue of the mammalian ala temporalis, and that the pro- 
cessus basipterygoideus is the homologue of the processus alaris 
of the ala temporalis. To explain the difference in the relations 
of the nervus maxillaris trigemini to the antipterygoid and ala 
temporalis, he assumes that the nerve has, in mammals, slipped 
over the top of the antipterygoid in early stages of development. 
I formerly concluded (14d) that the antipterygoid of La- 
certa was the homologue of the pedicel of the alisphenoid of 
Amia, and the processus basipterygoideus the homologue of the 
floor of the orbital opening of the myodome of Amia. The pars 
ascendens of the quadrate formed the lateral wall of the post- 
trigeminus portion of a trigemino-facialis chamber, as in the 
Amphibia. My present work leads me to consider these con- 
clusions correct, but to consider the trigemino-facialis chamber 
of these vertebrates to be the homologue of that chamber of 
Ceratodus and the Holostei, and not of the chamber of the Am- 
phibia and Teleostei; for the lateral wall of the chondrocranium, 
both of Lacerta and Crocodilus (Shiino, 714), is certainly the 
primitive cranial wall and not the outer wall of a trigemino- 
facialis recess. The processus basitrabecularis of Crocodilus 
would then represent a part of the floor of that chamber, and 
the processus pterygoideus quadrati a part of its lateral wall. 
The vena cardinalis anterior of Lacerta is said by Gaupp (00, 
pp. 547 and 548) to run posteriorly dorsal to the processus 
basipterygoideus and then along the external surface of the chon- 
drocranium, thus lying wholly external to that cranium. ‘This 
is exactly as it should be under my interpretation of the condi- 
tions, for this vein is the vena jugularis of my descriptions of 
