SEXUAL ACTIVITIES OF THE SQUID 401 
for three parts of what is here called the spermatophoric organ. 
The term does not serve my purpose, for the portion is more 
than a mere gland or indeed a series of glands. There are more 
parts to be decribed than the three divisions given by Marchand, 
and the term spermatophoric gland would indicate that the sper- 
matophores are formed here, while they are only partly formed 
here. 
The other portion of the mucilaginous gland (MG') forms a 
large outgrowth from the side of the portion just described. The 
opening of this portion is near the opening of the vas deferens 
and a considerable portion of the gland extends back between 
the viscera and the portion just described. This is called the 
first part of the spermatophoric gland by Marchand, and will 
be called the first part of the mucilaginous gland here. 
In structure the two parts are much alike. Both have thick 
walls, thrown into folds on the inside. These folds are fre- 
quently joined by bridges, and in many places the deeper por- 
tions of the depressions between the ridges form pouches or 
sometimes tubules (Williams ’08). The whole is, however, too 
open to form a true racemose or tubular gland. The cavity of 
each portion of the gland is extensive, forming a pelvis or basin in 
which the secretion is poured. The whole interior of the gland is 
ciliated, but the pelvis is particularly well supplied with cilia. 
The spermatozoa, entering from the vas deferens, pass into the 
pelvis of the second part of the mucilaginous gland (fig. 32), 
where they are mixed with secretion and the moving thread 
of sperm is covered with it. The spermatozoa do not enter part 
one of the mucilaginous gland, but are passed along a groove 
in part two past, but a little to one side of, the opening of part 
one. In the region of the groove, and for some distance along 
the side, especially along the side nearest part one, the cilia 
are large and numerous and serve to move the mixed sperm 
and secretion continuously toward and along this groove through 
this portion of the mucilaginous gland. 
Possibly one-third of the distance from the vas deferens to 
the distal end of part two the walls of the groove are thrown 
into a few spiral ridges (fig. 34, /’), between which the spermatozoa 
