440 WILLIAM M. GOLDSMITH 
Nowlin’s work (’06) on Coptocycla aurichalcea and C. guttata, 
though deficient in details, conforms with that of Stevens on 
thirty species, of which those considered above are types. In 
these forms Nowlin finds the unequal pair of heterochromosomes 
united, as usual, at metaphase; then they separate, pass to 
opposite poles, and divide normally in the second spermatocyte 
division. 
Miss Nichols (’10) finds that in Euchroma gigantea, ‘‘the 
small heterochromosome is separated from the larger in the first 
division.”’ Since she presents no further evidence and her 
figure 21 shows the two elements still united, her conclusion can 
have but little weight. The further meagerness of her observa- 
tions is evinced by the fact that neither the spermatogonial nor 
the second maturation divisions are figured. Even the number 
of chromosomes is not given. 
In Cicindela primeriana (’06) and C. vulgaris (09), according 
to Stevens, the heterochromosome group is trilobed in the meta- 
phase of the first maturation division. From this mass a small 
spherical chromosome separates and leaves a larger V-shaped 
one. Since each of these elements passes to opposite poles and 
divides normally in the second maturation division, the Cicin- 
delidae are placed with this type. The observations set forth in 
the present paper, however, do not substantiate these conclusions. 
It is observed that in each of the families Buprestidae, Ceram- 
bycidae, Chrysomelidae, Cicindelidae (according to Stevens, 
06, 09), Coccinellidae, Melandryidae, Meloidae, Scarabaeidae, 
and Staphylinidae an unequal pair of heterochromosomes occurs. 
The larger member of the pair is the maternal homologue of the 
odd-chromosomes in the Dytiscidae, Elateridae, and Lampyridae. 
The following is a typical fertilization formula for these 
families: 
Chelymorpha argus (Stevens, ’06) 
Sperm Egg 
(10+ Y)+ 00+X) =20+X4+Y=22¢ 
(10 +X) + (10+ X) = 200+ 2X £=22 9 
