CHROMOSOMES OF TIGER BEETLES 441 
Type II 
As was noted above, Doryphora decemlineata (Wieman, ‘10) 
contains, in the growth period, a conspicuous basic-staining 
bipartite body. This element takes its position on the spindle 
as in the thirty or more species reported by Stevens. However, 
instead of the two parts separating, the entire body passes to 
one pole and divides normally in the last maturation division, 
giving dimorphic spermatozoa with sixteen and eighteen chromo- 
somes. D. decemlineata is the only species of Coleoptera that 
has been recorded, prior to the appearance of the present paper, 
in which a bivalent sex-chromosome passes undivided to one 
pole. In this instance, however, neither the spermatogonial nor 
oogonial number is given. 
As will be shown in this paper, the double odd-chromosome 
in the Cicindelidae takes the characteristic position on the first 
maturation spindle and then, in contradiction to the records of 
Stevens, passes to one pole undivided. 
In the Carabidae, Hydrophilidae, Silphidae, and Tenebrion- 
idae some species have an odd-chromosome; others, an unequal 
pair of heterochromosomes. The Cicindelidae and at least one 
Hydrophilid (Wieman, ’10) possess a double odd-chromosome 
which passes to one pole of the first maturation spindle. 
The typical fertilization formula of the last-mentioned forms 
-in which the bivalent body passes undivided to one pole is as 
follows: 
Cicindelidae (present paper) 
Sperm Egg . 
104+ 104+X+x =204+ X+ x=22¢ 
10+X+x+(00+X +x) = 204 2X 4+ 2x = 24 9 
Type III 
The third type is characterized by the appearance of a single 
odd-chromosome. This type is represented by twelve species 
from seven families. Ten of these species were reported by 
Stevens. In all cases except two, the odd-chromosome passes 
undivided to one pole in the first maturation division and divides 
