460 WILLIAM M. GOLDSMITH 
that found in the male. In view of the fact that only two of 
the four female elements are visible, we might assume that the 
two large X’s have fused to form the large nucleolus, and the 
two small x’s to form the small one. The great variation, how- 
ever, in the size of these two bodies tends to weaken these 
assumptions. 
C. Bouquet, synizesis, and later stages 
It has been noted that in the typical condition the leptotene 
threads are scattered loosely throughout the nuclei and may be 
attached to the nuclear wall at any point. Immediately follow- 
ing this typical leptotene condition, the nuclear wall on the one 
side becomes free from leptotene threads (fig. 99). At this 
time more threads than usual are attached by only one end, the 
other end floating free in the cell sap. This free end soon finds 
its way to the ‘polarized’ side of the nucleus where the opposite 
end is usually attached. By this method the large loops of the 
bouquet stage are formed, and the nucleus is cleared on one side 
of chromatin fibers (fig. 100). This method also clearly accounts 
' for the fact that loops, rather than the ends of the leptotene 
threads, extend outward from the chromatin mass in the bouquet 
stage. As the loops are never drawn tightly together, there 
never appears a compact bouquet as described in other forms. 
Figure 101 is perhaps the most typical case found. No indica- 
tions of a pairing of these threads were observed. 
From the bouquet stage the threads emerge in broken pieces 
of more or less faintly stained chromatin rods (figs. 102 to 104). 
These appear very irregular and feathery, until the stage repre- 
sented in figure 105 is reached. In this and later stages the 
chromatin material is scattered uniformly throughout the large 
nucleus in the form of faint anastomosing aggregations. 
11. THE SOMATIC CELLS AND MITOSES 
The follicular tissue of the ovaries proved very satisfactory for 
a study of the somatic cells. Although the majority of the 
cells are in a resting condition, mitotic divisions are compara- 
