THE MEIOTIC PHENOMENA IN BLAPS 89 
A few V-shaped rods have their free ends directed toward the 
sphere, but this is inconstant. It is questionable whether their 
arrangement is merely accidental or constant for some euchromo- 
somes, the former assumption being most probable. Hence it is 
not possible to speak of a bouquet stage. This term was adopted 
in my first paper, but the comparison with forms in which this 
stage appears very marked shows that a great difference exists 
in this respect. . 
g. Pachytene stage proper. ‘The chromosomes in this stage 
appear spread throughout the nucleus without definite orienta- 
tion and are probably connected with each other by means of 
very delicate threads. As in the preceding stage, each euchro- 
mosome is a thick rod in which, at first, no split can be detected. 
In the late pachytene nuclei, however, a few chromosomes show 
the beginning of the longitudinal ‘split of the succeeding stage. 
The bivalent euchromosomes are continuous throughout; the 
chromatin granules do not show an arrangement into two series, 
which would suggest the existence of two conjugated 
chromosomes. 
h. Diplotene stage. As already mentioned, this stage lasts but 
a short time and immediately precedes the formation of the dense 
bivalents of the first maturation mitosis. It is questionable, 
therefore, whether we should consider such a condition as a sep- 
arate stage. Nothing can be added to the description already 
given in the preceding pages. 
C. The X-complex during the growth period. The X-complex is 
formed at the beginning of synizesis. Its components, sepa- 
rated up to this time and often scattered throughout the nucleus, 
come closer together and finally form a group which, from this 
moment, behaves as a single unit, moving to the pole nearest to 
the sphere along with the euchromosomes (fig. 71). At least the 
three large components can be detected and, in favorable cases, 
one or two smaller masses, which represent the small chromo- 
somes, are also found in close contact with the former. One of 
the large chromosomes appears somewhat denser than the others 
and retains this condition in the succeeding stages. This body 
is the X- or accessory chromosome. Its earlier history differs 
