190 EDWARD L. RICE 
inclines rather to the view that the active factor is to be found 
in the growth and expansion of the olfactory epithelium. Thus 
the extraconchal recess is primary and the concha secondary, 
the expansion of the nasal chamber being prevented at this point 
by the passive resistance of the gland. In a later publication 
(05 b) he modifies this view, and follows Born (’79, ’83), who 
accords even less of mechanical significance to the gland. The 
folding of the olfactory epithelium and its limiting cartilage 
give rise to the conchal invagination, “in die sekundiir die glan- 
dula nasalis lateralis von vorn her hineinwachst.’’ The develop- 
ment of Eumeces speaks strongly for the secondary relation of 
the gland to the concha. In stage 6 the gland lies, as described 
by Gaupp, deep in the aditus and the inverted trough of the 
concha itself. From this position the duct (figs. 27 and 28, 
d.l.na.) is easily followed along the lateral face of the cartilage 
wall of the zona annularis to enter the nasal cavity through 
the notch separating the superior alar process from the zona 
annularis. On the other hand, even as late as stage 5, the 
apparently solid anlage of the gland has grown backward only 
to a point slightly in front of the extreme anterior extension 
of the extraconchal recess, and thus well outside of the aditus 
conchae. The infolding of the concha is already well established, 
but the slowly developing gland rudiment is lagging far behind. 
In the lateral wall of the recessus extraconchalis of Lacerta 
(Gaupp figures an enormous fenestra lateralis nasi, upon which 
he lays some emphasis—‘‘ Auf die grosse Fenestra nasi lateralis 
sei aber besonders hingewilesen, mit Riicksicht auf die Kieferhéhle 
der Siuger, deren Entstehungsort in dieser Gegend zu suchen 
ist’”’ (00, p. 568). In Eumeces there is no suggestion of such a 
fenestra, the lateral wall being perfectly continuous in this region 
in stage 5 (figs. 25 and 26), and in all other observed stages in 
which the cartilage is recognizable; nor is there any weakening 
of the wall in the later stages to suggest an approaching resorp- 
tion of cartilage, as in the fenestration of the nasal and inter- 
orbital septum described above. 
Gaupp (’05b) has himself called attention to the recorded 
absence of the fenestra lateralis in the crocodile (Parker, ’83) 
