380 MITCHEL CARROLL 
950 and 2482, it seems to have the same history as in Pentatoma 
and Rhomaleum, appearing in the growth stages as a large body 
staining like chromatin and lying in the cytoplasm, though often 
pressed against the nuclear membrane. In individuals of this 
type it is usually a conspicuous body also at the first matura- 
tion division (c¢ in figs. 46 and 48), but smaller similar staining 
granules may be present in the same cells (c in figs. 46 and 48). 
The latter I have called ‘chromatoid granules.’ One at least of 
these granules, which are sometimes seen in the cytoplasm, may 
be intranuclear in origin; for a small chromatic spheroid is 
often seen within the nucleus at diakinesis, while the nuclear 
membrane is still intact. A possible source of this spheroid may 
be seen at c, figure 47, plate 5, where a granule or ‘chromomere 
vesicle’ (Carothers, °17) is connected with a spermatogonial 
chromosome by a thin strand. The rest, judging from condi- 
tions observed in other individuals, may be homologues of the 
larger body. The large body has never been seen to divide 
(though I have observed it in a few instances apparently breaking 
up at the first spermatocyte kinesis). It usually remains pas- 
sive in the cytoplasm, and consequently at the end of the first 
mitosis is found in but one-half of the second spermatocytes. 
It starts to disintegrate, usually in the second spermatocytes, and 
when seen during the second mitosis is much smaller than in the 
first. Occasionally, however, it may be as large even in the sper- 
matids as in the first spermatocytes. The substance of the body 
seems to pass into the tail of the developing spermatozoon and 
to be extruded as described by Wilson and Plough. 
Rarely in the above individuals two nearly equal chromatoid 
bodies can be seen in the same first spermatocyte. 
In individual 954.1 a conspicuous chromatoid body is present 
in the growth stages, but almost invariably fragments before the 
first maturation division. In a few cells (fig. 15, c) a small chro- 
matic body was observed in the metaphase, in addition to the 
usual chromosomes. This element lies in the spindle and appears 
more like a chromosome fragment than a chromatoid body. It 
is too small to be identified with the extra chromosome to be 
described presently. 
