504 A. B. DAWSON 
wide distribution and relatively high state of development make 
it impossible for one to regard them as unimportant in the life 
of the animal. They may never discharge, but, 1) may serve 
as reservoirs for a poisonous secretion (Cohn, ’95), or, 2) they 
may produce intercellular secretions of value in keeping the 
skin in a healthy condition (Pfitzner, ’79; Prowazek, ’01), or 
3) they may actively assist in molting, or, 4) they may discharge 
at the time of molting, producing a secretion which protects 
the naked cells of the replacing layer until a cuticular margin 
has been developed. Satisfactory evidence as to which of these 
views is correct is still wanting. 
E. Goblet cells 
Goblet cells were found only in the region of the mouth and 
within the gular fold. Kingsbury (’94) states: ‘The transition 
from the outer skin is gradual; the glands of the cutis and the 
cells of Leydig cease upon the outer side of the lip, while the 
first goblet cells appear within the line of teeth.” I, however, 
have found many goblet cells outside of the line of teeth, that 
is, on the lips. 
In highly developed cases (fig. 16) the goblet cell is sian 
egg-shaped and tapers at the top into a narrow neck, opening 
between the cuticular cells. Its body has a reticulated structure 
and stains lightly. A much flattened nucleus surrounded by a 
small portion of non-metamorphosed cytoplasm lies at the base 
of the cell. 
In the gular fold the goblet cells are similar to those of the 
mouth. I have been able to find only one reference to a similar 
local development of goblet cells. For axolotl, Paulicki (785) 
reports goblet cells on “der inneren Fliche des Kiemendeckels.” 
F, Pigment 
1. Distribution. The epidermis is deeply pigmented on the 
dorsal surface of the animal. Here the density of the epidermal 
pigmentation is quite closely correlated with that of the pigmen- 
tation of the adjoining dermis. Very little pigment is found in 
