28o Journal of Comparative Neurology. 



and these rami are somewhat differently composed. It receives, 

 as in Menidia, the ventral division of the dorsal lateralis root 

 (r. buccalis), the whole of the motor V root, the larger part of 

 the general cutaneous fibers from the Gasserian ganglion and a 

 considerable number of communis fibers from the geniculate 

 ganglion. The motor V fibers run out in the midst of the 

 sensory V, from which they can be distinguished by their larger 

 caliber. The r. buccalis occupies the dorsal part of this com- 

 plex and the communis component the ventral. The relations 

 of the components as the trunk passes through its foramen are 

 shown in Fig. 7. 



Just outside of the foramen the trunk divides into three 

 approximately equal divisions, the r. maxillaris, the r. mandi- 

 bularis V and the r. buccalis. The motor V fibers clearly go 

 out with the r. mandibularis. The communis fibers from the 

 geniculate ganglion could not be so clearly followed, as they 

 divide up into several fascicles which interlace with the general 

 cutaneous fibers from the Gasserian ganglion. Communis fibers 

 can, however, be seen to enter both the r. maxillaris and the 

 r. mandibularis V. 



The r. buccalis. — I have followed this nerve for its entire 

 length in my sections, but can add little to Cole's account 

 of it. The otic and outer buccal nerves are, as stated above, 

 accompanied by general cutaneous fibers, corresponding, as 

 Cole has mentioned, to nerves in the same relation in the tad- 

 pole of the frog (Strong, '95) and in Amia (AUis, '97). One 

 of these twigs was traced forward onto the cornea. A minute 

 twig is given off from the outer buccal nerve between the 7th 

 and 8th infra-orbital canal organs for naked pit-organs ventrally 

 of the latter organ (Fig. i, b. p. 0.). These organs lie caudad 

 and dorsad of those supplied by the branch H^ of the hyoman- 

 dibular trunk. 



I confirm Cole's account of the course of the inner buccal 

 branch completely ; the only differences which I find are in the 

 branches to the pit-organs about the infra-orbital canal and 

 these slight differences do not seem to be of any morphological 

 importance, but rather matters of individual variation. Most 



