300 Journal of Comparative Neurology. 



particularly the prc-vagal sympathetic and the r. lateralis acces- 

 sorius. Let us take up the analysis of the trigemino-facial 

 complex as given on p. 133 of his paper. 



(i) His lateral line ganglion requires no further comment. 

 My findings agree with his description. 



(2) His " trigeminal or Gasserian " ganglion "is situated 

 internal and largely anterior to the second or ventral portion 

 of (i). From this portion of the complex the superficial oph- 

 thalmic of the trigeminus, the maxillo-mandibular trunk, and, 

 probably, the post-branchial division of the facial arise. If the 

 latter statement be correct, as seems certain, this ganglion 

 should be called the ' trigemino-facial.' " The last supposition 

 proves to be correct. We have two quite distinct ganglia here, 

 the Gasserian and the geniculate or facial, as my descriptions in 

 the preceding part have shown. If these two ganglia are clearly 

 distinguished (which Cole failed to do) the difficulty, to which 

 the author refers in the foot-note, in the interpretation of the 

 morphology of the proper Gasserian ganglion of Gadus, does 

 not exist. The case is perfectly simple ; the Gasserian ganglion 

 is strictly homologous with the ganglion of that name in Me- 

 nidia, Rana, Amblystoma and all other vertebrates in which it 

 has been clearly distinguished from the other ganglia of the 

 complex. From the fishes to man this ganglion gives rise to 

 all of the pre-auditory general cutaneous fibers and, so far as 

 now known, to no others. 



(3) His " facial " ganglion is thus described: " This is a 

 long and perfectly distinct ganglion which lies ventral to (2). 

 It is connected with the palatine nerves and also with the pre- 

 spiracular or chorda tympani divisions of the facial. It also 

 communicates (a) directly with the sympathetic trunk ; (d) by 

 means of Jacobson's anastomosis (see below) with the glosso- 

 pharyngeus." This description was most puzzling to me until 

 I discovered that the "facial" ganglion here referred to is in 

 reality the sympathetic ganglion. The more detailed account 

 of this ganglion which he gives on p. 135 leaves no doubt 

 of this. 



How, then, shall we account for the fact that the "facial 



