3o6 Journal of Comparative Neurology. 



passes upwards, receives a root from the vagus, and then doubt- 

 less has the usual peripheral distribution." 



Haller seems to regard this accessorius of the carp as a 

 general cutaneous nerve contributing to the brachial plexus 

 for the region of the pectoral fin. Cole regards it as a "so- 

 matic sensory," i. e., cutaneous, nerve from the Gasserian gan- 

 glion, which "collects" the dorsal branch of the "hypo- 

 glossal," and a vagal branch and then proceeds into the trunk 

 to form the accessory lateral line nerve, there to collect the 

 dorsal branches of the spinals. My own view of the accessorius 

 of cyprinoids, as developed in Section 12 of the Menidia paper, 

 is different from either, I have regarded it as an intra-cranial 

 anastomosis between the geniculate ganglion and the vagal root 

 of the r. accessorius. It should be noted that Haller's account 

 is based on dissection only. He has made no accurate analysis 

 of the trigemino-facial ganglia and there is no satisfactory evi- 

 dence that the root in question arises from the trigeminal, or 

 Gasserian ganglion, rather than from the facial, or geniculate 

 ganglion. Haller does not state that the nerve arises in part 

 from the facial ganglion, but from the " Facialisaste des Tri- 

 geminus," and his trigeminus ganglion, judging from his figures, 

 is quite as likely to be the geniculate. 



Stannius' description ('49, p. 60) of the conditions of this 

 root in the cyprinoids will prove, I think, much more accurate: 

 "Was die Quelle des Ramus recurrens anbetrifft, so ist derselbe 

 von der dritten Wurzel des Nervencomplexes, niimlich derjeni- 

 gen, welche aus dem Lobus impar medullae oblongatae ihren 

 Ursprung nimmt, abzuleiten. Diese starke Wurzel, welche 

 ausschliesslich feine Primitivrohren enthalt und motorischer 

 Eigenschaften ermangelt, spaltet sich noch innerhalb der Sched- 

 elhohle in mehre, schwer zu trennende und zu verfolgende 

 Strange. Einer derselben tritt iiber an den eigentlichen N. tri- 

 geminus und ist namentlich in den an der Austrittsstelle des- 

 selben gelegenen gangliosen Plexus zu erfolgen, gibt auch- 

 Elemente ab zu den in der Schedelhohle aufsteigenden Nerven. 

 Ein zweiter Strang hilft den N. facialis zusammensetzen. Ein 

 drittes, betrachtlicheres Biindel tritt iiber in eine umfangliche 



