310 Journal of Comparative Neurology. 



dorsal series continuing the main line of the head and in a ven- 

 tral series under the pectoral fin. No terminal buds are de- 

 scribed along- the course of this nerve. The condition in Pro- 

 topterus (Pinkus, '94) is probably essentially similar, though 

 perhaps more extreme, for here the nerve r. lat. c. VII + X 

 of Pinkus' figures, which might be interpreted as the facial root 

 of the r. lateralis accessorius, is described as wholly composed 

 of lateralis fibers. 



As already stated in the descriptive part, I have not been 

 able to demonstrate any general cutaneous fibers entering the 

 r. lateralis accessorius either from the Gasserian ganglion or 

 from the jugular ganglion. Nevertheless the conditions are 

 such that it is impossible to exclude the possibility that a small 

 number of such fibers may enter this nerve from one or both of 

 these sources. In any case we are certain that branches of the 

 r. lateralis accessorius effect terminal anastomoses with branches 

 of the r. cutaneous dorsalis vagi, as well as with dorsal rami 

 of spinal nerves farther back. Miss Clapp's figure ('99, PI. 

 -XVIII, fig. 13) suggests that the ramus lateralis accessorius in 

 Batrachus receives important general cutaneous accessions from 

 both facial and vagus roots (especially clearly in the latter case), 

 and this again can be correlated with Strong's discovery in the 

 tadpole of the frog of an extra-cranial general cutaneous anas- 

 tomosis from the vagus to the facialis. The condition which 

 he figures and that figured by Dr. Clapp could both be derived 

 from a case in which the r. cutaneous dorsalis vagi effected a 

 broad extra-cranial anastomosis with the trunk of the r. later- 

 alis accessorius. In Batrachus the communis fibers which pri- 

 marily composed the r. lateralis accessorius appear to have been 

 largely replaced by lateralis fibers (and probably general cuta- 

 neous fibers also). The extra-cranial anastomosis with the 

 vagus may represent the vagal root of the r. lateralis accessor- 

 ius ; but if so its communis fibers have joined the r. cutaneous 

 dorsalis vagi, for the origin of this anastomosis from an intra- 

 cranial vagal ganglion and the course of its fibers beyond the 

 anastomosis with the accessorius (as described to me personally 

 by Miss Clapp) show unmistakably that this nerve is chiefly the 



