George L. Streeter 161 
taste fibers. Reasoning from the development and comparative anatomy 
of the chorda tympani he points out the improbability of the presence 
of any taste fibers in the trigeminal nerve as had been maintained by 
many. That the fifth nerve pair contains few or no taste fibers has been 
further established by the careful observations of Cushing, 04, who, after 
the removal of the Gasserian ganglion, fund in all of his cases that 
the anterior part of the tongue retained in greater or less degree the 
sense of taste as well as common sensation. 
If after the work of Sapolini there remained still doubt regarding 
the anatomical identity of the pars intermedius, geniculate ganglion, and 
chorda tympani, it was removed by the tonvincing dissections of Penso, 
93. This worker investigated these structures in man and a considerable 
variety of other mammals, using principally dissection and teasing 
methods, though his observations were made also in part from serial 
auric. vag! 
iz 
\ 
| 
' 
\ 
| 
| 
! } : 
gang. genic. 
petros. sup. maj. 
\- -- -motor VW. 
n.chord. tymp. 
Fig. 7. Drawing made from a dissection of the facial nerve in a 20 cm. 
pig embryo. At this stage the sensory and motor divisions can be easily 
separated. 
sections. He showed conclusively that.the geniculate ganglion is primar- 
ily connected with the motor trunk of the facial only by contiguity, its 
fundamental attachments being a central one, the pars intermedius, and 
two peripheral ones, the chorda tympani and the great superficial petrosal. 
He also describes a number of small anastomoses existing between these 
branches and the surrounding structures, including the motor portion 
of the facial, the acoustic ganglion, the spheno-palatine ganglion, and 
the auricular branch of the vagus. He therefore represents the great 
superficial petrosal and chorda tympani as composite nerves, which are 
made up, in the first place of fibers from the geniculate ganglion, and 
in addition to these the fibers from the above-mentioned anastomosing 
branches. During the past year a paper confirming Penso’s work in its 
