SACRAL SYMPATHETIC TRUNK OF CAT 87 
bodies. Huber states further that the fibers thus ending did 
not seem to enter the ganglion from the periphery, i.e., did not 
seem to be terminations of neuraxes whose cell bodies were 
situated distal to the ganglion under observation. Huber’s later 
observations (’99, ’13) do not appear to alter his position with 
reference to these fibers. 
More far-reaching are the observations of Dogiel, who in 
1896 described two types of sympathetic cells, 1) the commonly 
recognized postganglionic motor neuron with cell-body located 
in a sympathetic ganglion and, 2) another type of cell with 
usually a larger cell body and much longer dendrites. The 
dendrites branch less and were often traced beyond the limits of 
the ganglion. The neuraxes of these cells were sometimes 
traced into another ganglion, where they were seen to divide 
repeatedly and take part in the formation of the intercellular 
plexus. Dogiel expresses the belief that these cells are sympa- 
thetic and of sensory function. He refers further to the proba- 
bility that the pericellular baskets about spinal ganglion cells of 
type II take their origin from sensory sympathetic cells. 
While the observations of the three authors last mentioned 
have not been materially extended or substantiated by subse- 
quent investigations, it is also true that anatomical and physio- 
logical evidence is hardly sufficient as yet to rule out the possi- 
bility of the existence of sympathetic commissural neurons. 
The writer’s contribution to this phase of the structure of the 
sympathetic trunk is based on an experimental study of the 
sacral and anterior coccygeal parts of the sympathetic trunk 
of the domestic cat. As stated previously, this part of the 
trunk in the cat is especially favorable, as it is connected with 
the spinal nerves only by gray rami communicantes, the lowest 
white ramus being that of the fourth or the fifth lumbar nerve 
(Langley, 92). No preganglionic efferent fibers run from the 
spinal cord to the sympathetic through the gray rami (Langley, 
96). It follows that all preganglionic efferent fibers in the sacral 
trunk must run through the lower thoracic and lumbar white 
rami and descend in the trunk to the sacral and coccygeal region. 
Division of the trunks caudad to the lowest white rami should 
