1912] Lacinia in Maxilla of Hymenoptera 235 
‘Galea. The galea (we incline to believe this part homolo- 
gous with the galea of the locust’s maxilla, rather than with the 
lacinia, because of its two-segmented condition) extends distad 
from the stipes as a tapering blade-shaped piece. It is composed 
of two segments. The proximal one is small and triangular, 
articulating by the entire length of one of its margins with the 
stipes. The distal segment or sclerite constitutes the real 
blade-like portion of the maxilla, and nearly equals in length 
the ligula and labial palpi (see below). Its surface is unequally 
divided into two portions by a submedian, dark-brown, longi- 
tudinal line. (This line may indicate a coalescence of galea and 
lacinia into this one blade-like compound sclerite). This line 
bears several hairs, and there are scattering hairs elsewhere on 
the sclerite, especially toward the distal end.”’ 
Snodgrass} writes as follows of the maxilla of the honey-bee: 
‘‘Let us now return to a study of figure 15D. The series of 
lateral pieces as already explained are the maxilla. A com- 
parison with figure 3B representing a generalized maxilla will 
show that these organs in the bee have suffered a greater modi- 
fication than has the labium, but the parts can yet be quite 
easily made out. The main basal plate (st) is the combined 
stipes, subgalea, and palpifer, the basal stalk is the cardo (cd), 
and the little peg-like process (mx plp) at the outer end of the 
stipes is the greatly reduced maxillary palpus. Hence, we have 
left only the terminal blade-like lobe (mx) to account for, and 
it is evident it must be either the galea or the lacinia (See fig. 
3B, ga and Jc) or these two lobes combined. Here’ again a 
comparative knowledge of the mouth parts of Hymenoptera 
comes to our aid and shows clearly that the part in question is 
the outer lobe or galea, for the inner one becomes smaller and 
smaller in the higher members of the order and finally disappears.” 
There is expressed in these two quotations very different 
views, the former that the galea and lacinia are probably 
coalesced and the latter that the lacinia is wanting. This is 
the status of the lacinia in the higher Hymenoptera, writers 
consider it either as fused with the galea or as obsolete. 
A comparison of the drawing of the maxilla of the honey 
bee with that of Bombus shows it to be similar in form but 
shorter and consists of a long, slender, proximal piece, the cardo, 
Snodgrass, R. E.—The anatomy of the honey bee. U.S. Dept. Agr., Bur. 
Entom., Tech. Ser. No. 18, 1910. Pp. 45-46. 
