Anatomy of the Male Genital Tube in Coleoptera. 588 
Heliocopris mouhotus. 
Median lobe small, entirely hidden and embraced by the lateral 
lobes. Lateral lobes rather large, with a membranous connection 
to near their tips, which membrane folds together when the lateral 
lobes approximate. Basal-piece large, forming an irregular chitinous 
tube, curved and enlarged at the base, with a large basal opening. 
Internal sac large, with two blunt, flattened, chitinous spines near 
apex. 
Onthophagus fracticornis. 
Median lobe small, with small chitinous support at base projecting 
into basal-piece as two short, broad, rounded, median processes (cf. 
Oniticellus). Lateral lobes small, connected by membranes to near 
their tips. Basal-piece large, forming chitinous tube with large basal 
opening at base. Internal sac large, bearing complex, curved 
chitinous plates on the apical half. 
Oniticellius marsyas. 
Tambour cylindrical, basal portion short. Lateral lobes short 
and powerful, abruptly flexed, of irregular, complicated form ; their 
median aspects contiguous throughout. Median lobe entirely 
concealed, forming at the base a chitinised tray, basally split for 
two-fifths of its length and forming a secondary tambour within 
the normal one. Sac largely developed. This is a very remarkable 
and high form of Coprinae, though the affinity with Onthophagus is 
a close one. 
Obs.—The Coprinae have the basal-piece rather com- 
pletely tubular in form, owing to the shortening of the 
basal part of the “tambour.” The basal part of the 
tambour (or great basal sclerite of the Scarabaeidae) is 
chitinous on one aspect, membranous on the other, and 
this basal portion being in Coprinae of small elongation in 
comparison with the distal portion, the tubular form of 
the distal portion is unusually conspicuous. The aedeagus 
of Coprinae is easy of recognition. 
Aphodius punctato-sulcatus (Pl. XLV figs. 18 and 18a). 
Median lobe small and membranous, supported on the dorsal edges 
by two chitin strips (a) which project into the basal-piece as two struts 
(ms) for the support of muscles ; median orifice occupying all the 
dorsal face. Lateral lobes large, a semi-chitinous extension along 
the dorsal edges (b) form two flanges which overlap and hide the 
median lobe. Basal-piece large, the distal half forming a tube, the 
