Anatomy of the Male Genital Tube in Coleoptera. 617 
may, from our point of view, be more correctly placed in 
Trogositidae. Thymalus and Leperina depart from the 
more typical Trogositidae by the lateral lobes being 
ventrally brought together (completely conjoined in 
Thymalus, incompletely in Leperina). This point is of 
importance, because on account of it we have associated 
with the Cucujoidea certain families that have been 
usually associated in Heteromera. The tubular sheath 
formed by the tegmen in Trogositidae, is found in Cleridae, 
Byturidae, and in a somewhat different form in Cyatho- 
ceridae, and we have therefore placed the families in 
question in the Cucujid-Trogositid complex. 
The curious genus Diagrypnodes of Cucujidae will 
have to be separated from the family; it approaches 
Pythidae. On the other hand no surprise will be felt at 
the association of Pythidae and Aegialitidae (which are 
pretty certainly but one family) with Cucujidae, when it 
is recollected that the Cucujidae include Heteromerous 
forms, and that certain genera, e.g. Rhinomalus and 
Hemipeplus, have for long been sources of perplexity, as 
to the distinctions between “ Heteromera” and Cucujidae. 
Anthicus, Heteroceridae, Othniidae and Lathridius have 
but little specialisation of the sac; none of them show 
any special approximation to Cucujidae, but they appear 
to be less ill-placed in Cucujoidea than elsewhere. Lathri- 
dius is usually placed in one family with Corticaria, but 
the two have but little connection, and Corticaria will 
perhaps find a better position near Cryptophagidae, though 
it appears to be very aberrant. 
We have no hesitation in placing Coccinellidae in this 
complex, although Verhoeff (am Arch. Naturges, 61, 1, 
1895) has separated Coccinellidae as the equivalent of all 
other Coleoptera by the nature of the male structures, 
they possessing, according to his perception, within the 
“penis” (= our median lobe) a structure he calls the 
siphon. We do not take the same view of the structures 
as Verhoeff does. According to our view the siphon is 
the median lobe (penis of Verhoeff) and the part that 
hoods it (and that Verhoeff calls penis) is an unusual fold 
which is certainly a part of the tegmen, though we do not 
feel called on to decide as to its exact nature without a 
knowledge of the ontogeny. If this view of the structures 
be correct, Verhoeff’s two divisions of Coleoptera, viz. 
Siphonophora (= Coccinellidae) and Asiphona (= all other 
