Anatomy of the Male Genital Tube in Coleoptera. 625 
of the families combined with Tenebrionidae to form the 
“ Heteromera” must be separated. The comparatively 
small families, Cistelidae (Alleculidae of many recent 
writers), Lagriidae and Rhysopaussidae, are really allied 
to the huge group Tenebrionidae. Of this latter complex 
it will be noticed that we have examined but few forms. 
Taking Pediris as a central one we find the tegmen con- 
sisting of an elongate tubular basal-piece, chitinous on 
the dorsal aspect, with well marked and separate lateral 
lobes, making lateral and dorsal protection of the elongate 
median lobe. In the more specialised Tenebrionidae (e.g. 
Eleodes and blaps) the lateral lobes are soldered together, 
and the median lobe is reduced in extent. In certain 
cases (Cossyphus) the median lobe is reduced to a nearly 
or quite membranous condition, and in Cistelidae and 
Lagriidae there is a similar reduction. In the type of 
the Nosoderma-group we have examined (Zopherosis) there 
exists a distinction from Pediris that we must treat as of 
considerable importance, inasmuch as the chitinisation of 
the tegmen occurs on the ventral aspect, the lateral lobes 
being united in that position. This suggests that a com- 
plete sundering of the Tenebrionidae will be found neces- 
sary. Stenosis agrees with Zopherosis in this respect. It 
would be well worth examining Adelostoma and allies to 
ascertain whether there is a real affinity between the 
Stenosis and Zopherosis forms, but we have not been able to 
carry our investigations of the Tenebrionid forms farther 
than the inadequate extent that will be found in our 
anatomical section. 
As regards the families of “Heteromera” other than 
those mentioned above, we have already said that we 
have failed at present to connect them with the Tenebri- 
onidae; and we have assigned them tentatively positions 
in the Cucujoidea. Whether the Tenebrionidae really 
link on as further differentiations thereof (cf. Melandryidae 
and Pythidae) we are not prepared to express an opinion. 
We have left Monommidae in the Tenebrionid division 
(along with the Stenosis-Zopherosis forms), but it appears 
to be really very isolated. 
We may conclude our brief remarks on the Tenebri- 
onidae alliance by referring the reader to what we have 
said elsewhere as to the “simple trilobed form of aedeagus,” 
and adding that there is not here a great departure 
therefrom. 
