1917] Phylogeny of the ElateridcE 259 



CARDIOPHORINyE. 



The subfamily Cardiophorinas includes the single tribe 

 Cardiophorini and is characterized in the larval condition* 

 by the membranous integument of the body, by the accessory 

 digitate anal lobes, by the mandibles bearing teeth upon the 

 exterior surface, and by the spiracles being placed upon retractile 

 papillae. The adults are easily distinguished by the truncate 

 prosternal spine and the cordiform scutellum. 



Addenda. 



The placing of the following tribes in this classification is 

 merely suggestive as no larvae are known from any species con- 

 tained therein and my conclusions are based entirely upon adult 

 characters. 



The first of these tribes, the Oxynopterini, is composed of 

 four genera, and is characterized by the very excavate front, 

 protuberant simple mandibles, simple tarsi, very elongate 

 maxillary palpi, and the extremely broad metathoracic epimera. 

 Hope separated this tribe from the other Elateridse, erecting 

 therefor a family, the Phyllophoridae. The tribe seems to bear 

 affinities with Campsosternus and Tetralohus. 



The second, the Tetralobini, consists of the remarkable 

 African Elaterids of the genus Tetralohus. Enormous insects 

 measuring from 30-80 mm. in length, and characterized by the 

 "plaque nasale" being as broad as long; mandibles dentate; 

 antenna strongly flabellate in male; and the tarsi lobed. They 

 are probably closely related to the Oxynopterini. 



The third unplaced tribe, the Eudactylini, which Candeze 

 places with much reservation near the Dicrepidiini, is char- 

 acterized by having the frontal carina well defined; the pro- 

 sternal sutures fine and straight; tarsal joints short, very much 

 dilated and often lamellate and the claws simple. 



The fourth tribe, Crepidomini, lacks the frontal carina, the 

 front flat or concave and usually acuminate; posterior coxae 

 gradually narrowed inwardly; and joints two to four of tarsi 

 dilated. The tribe is confined to the Australian region and is 

 allied to Ludiina and Lepturoidina. 



* Figured in Bureau of Ent. Bull. n. s. 156, p. 8, Fig. 3b, 1915, and in greater 

 detail in Proc. Ent. Soc. Wash. XVII, p. 179-185, PI. 20, 21, 1915. 



