191' 



Phytogeny of the Elateridce 



263 



such in our present genus Elater is hardly justified. It is 

 probably safe to place the original Elaterid stem in the upper 

 Jurassic. The palontological record is blank from the time of 

 these earlier fossils until the early Tertiary. The baltic ambers 

 of the Lower Oligocene contain Elateridae so well differentiated 

 as to be referable to extant genera. Eucnemidae are very 

 numerous, as would naturally be expected in Amber faunas. 

 The Cardiophorinae had already split off from the parent stock, 

 probably in the Upper Cretaceous. The Pyrophorinse were 

 already differentiated into their main tribes, but what seems 

 most remarkable is that the Lepturoidini and not the more 

 conspicuous Pyrophorini are represented. In fact, the Pyro- 

 phorini are not recorded until the Upper Miocene where they 

 abound in both the Oeningen beds in Europe and in our own 

 Florissant of Colorado. This seems to indicate that our tribe 

 P^^rophorini left the Pyrophorinas stem at a much later period 

 than the tribe Lepturoidini. The great gap in the record 

 covers the period in which the greatest Elaterid diversification 

 took place, that is in the Cretaceous and lower Eocene. 



Family ELATERIDAE. 



Italicized tribes and sub-tribes are tentatively placed, as larvae of species 

 have never been described. 



Subfamily Pyrophorinae 



1. Tribe Pyrophorini 



Sub tribe Agrypnina 

 Sub tribe Hemirhipina 

 Sub tribe Chalcolepidina 

 Sub tribe Pyrophorina 

 Sub tribe Monocrepidina 



Tribe Oxynopterini 



Tribe Tetralobini 



2. Tribe Pityobini 



Sub tribe Pitj'obina 



3. Tribe Lepturoidini 



Sub tribe Athouina 

 Sub tribe Lepturoidina 

 Sub tribe Ludiina 

 Tribe Crepidomini 



4. Tribe Oestodini 



Sub tribe Oestodina 



Tribes not yet located: 

 Tribe Allotriini 

 Tribe Dimitini 



Subfamily Elaterinae 



1. Tribe Steatoderini 



Sub tribe Sericosonina 

 Sub tribe Steatoderina 



2. Tribe Agriotini 



Sub tribe Agriotina 

 Sub tribe Adrastina 



3. Tribe Elaterini 



Sub tribe Elaterina 

 Sub tribe Dicrepidina 

 Tribe Eudactylini 



4. Tribe Physorrhini 



Sub tribe Physorrhina 



5. Tribe Melanotini 



Sub tribe Melanotina 

 Subfamily Cardiopherinae 

 Tribe Cardiophorini 



Sub tribe Cardiopherina 

 Subfamily Physodactylinae 



Tribe Hypodesini 

 Tribe Cardiorhini 



