The Lateral Nasal Glands of Amphiuma. 151 



take the paths of least resistance, and would naturally work their way 

 along the larger nerve bundles. A more probable source of innerva- 

 tion of the gland is from the ramus glandularis opthalmici profundi I, 

 the branches of which spread out in the connective tissue of the 

 snout dorsal and lateral to the gland; some of these fibers may be 

 traced to the skin of the snout, but several intermingle with the 

 anterior portion of the gland and seem to lose themselves within its 

 mass.^ 



II. The Development of the Lateral Glands. 



As the accompanying table shows, my study of the development of 

 the lateral gland has been based upon serial sections of six specimens 

 ranging in total length from 30 mm. to 190 mm. To this list may 

 be added a single specimen 80 mm. in length in which the glands 

 were studied by dissection. This latter method, while not abso- 

 lutely trustworthy as to the exact relationships of the individual 

 tubules, furnishes an extremely satisfactory corroboration of the 

 results obtained by the more laborious method of reconstruction from 

 serial sections. The dissections were made 'by slicing off with a 

 sharp scalpel the lateral portion of the head of a specimen well hard- 

 ened in alcohol, the cut passing obliquely through the nasal cavity 

 in such a way that the whole of the external naris was included in 

 the portion removed. This detached portion was then pinned out 

 in a small dissecting pan and the work of exposing the gland was 

 completed by dissecting from within the nasal cavity, since the only 

 parts to be removed were the nasal epithelium and the cartilaginous 

 capsule. 



In the smallest specimen sectioned, a larva 30 mm. long, there is 

 no trace of a lateral nasal gland or of nasal muscles. 



The next stage, a 60 mm. specimen in adult form, shows a single 

 tubular gland (Fig. 5 a) upon each side. Towards the posterior end, 

 the lumen of the tubule becomes much enlarged and ultimately divides 

 into two lumina, although in the external wall there is no evidence 



*An article by Norris, '08, on the Cranial Nerves of Amphiuma has appeared 

 since this manuscript left our hands, in which it is stated that neither of 

 these branches innervate the lateral gland, and that the name ramus glandu- 

 laris is therefore a misnomer. 



