502 Howard Edwin Enders. 



to its anterior end nsiially pass into the mouth by a peculiar mus- 

 cular contraction. As they reach the anterior end of the ciliary 

 groove the dorsal lip of the buccal funnel is drawn backwards and 

 the ciliary groove, which now extends beyond the dorsal border of 

 the mouth, permits the granules to fall directly upon the ventral 

 lip of the funnel. The dorsal lip is then pushed forward and takes 

 its normal form. Other granules pass from the anterior end of the 

 ciliary groove laterally, along the posterior margin of the dorsal 

 lip of the funnel, l^ear the mesial side of the tentacles some of the 

 granules are swept over the dorsal lip and into the funnel, others, 

 however, continue laterally posterior to the tentacles and the dorsally- 

 directed auriculate lobes of the ventral lip. They are discarded at 

 the side of the animal in shreds of mucus. Some particles of car- 

 mine which had traversed the mesial ciliary groove to its anterior 

 end were swept to the right and left of the groove and there were 

 swept backwards in two parallel rows to the vicinity of the twelfth 

 segment. Here they were either discarded at the sides of the animal 

 in shreds of mucus or were again swept into the forward current 

 and, thence to the mouth, where they were disposed of as described 

 before. 



The particles which dropped upon the ventral lip were swept, 

 indifferently, into the esophagus or towards the margin of the lip 

 and discarded. The statement has been made in another part of 

 this paper that in the accessory feeding organ of the thirteenth seg- 

 ment the granules of carmine are swept from its margin into the 

 deeper portion where they are rotated and mixed with mucus. The 

 accessory feeding organ is tilted so far forwards that it overhangs 

 the mesial ciliary groove when the thirteenth segment, which bears 

 it, is contracted (Figs. 3 and 4). Its muscular lips open widely and 

 the mass of accumulated material is pressed out by contraction and 

 flattening of the whole pouch. The carmine boluses Avere several 

 times dropped on the mesial ciliary groove. They were then swept 

 forwards to the buccal funnel, but were in each case discarded. One 

 small mass of diatoms Avhich was thus discarded from the accessory 

 feeding organ was swept into the esophagus after it had traversed 

 the mid-dorsal ciliary groove and was dropped on the strong cilia 



