109 
ANTHESIS DATES 
MAY JUNE JULY 
1973 oO O &—A Aa 
Alopecurus = es BP ea as ae = — eee ee ES —— 
alpinus 
1974 o—oO O a 4A 
1973 OS 
Deschampsia wets 
caespitosa wa 4 =x = _ a i i ae 
1974 O—O @—@ Oa/ 
1973 2 eeeeaecrens "| A 
Calamagrostis ie eee ee. O O 
inexpansa ae St 1S a Nee pike 
4 1974 o—_) Aa/ 
QL 0 
1973 Se A 
Festuca fone WE pa ® 
rubra a ie eee a eae cs aw =p, = 
1974 ——r A a/ 
O Arctic biotypes 
O Boreal, forested biotypes 
® Boreal, alpine biotypes 
LZ Boreal, coastal biotypes 
A Colorado alpine biotypes 
9/Anthesis occurred after June 20; specific dates not obtained after that date in 1974. 
Fig. 19. Anthesis dates of transplant entries in Palmer garden in 1972 and 
1973 organized according to type of origin. 
highest polyploid in Ca/amagrostis inexpansa 
grew the most at Palmer but failed at Prudhoe. 
Extensive sampling and testing at the population 
level would be necessary to better define dif- 
ferences related to ploidy levels. 
A comprehensive transplant study involving 
populations of Agrostis tenuis led Bradshaw 
(1959) to emphasize the roll of small popula- 
tions interacting with their particular environ- 
ment to produce local differentiation. He found 
natural selection effective in distinguishing 
populations occupying habitats very short dis- 
tances apart. Though conditions in the North, 
particularly in the Arctic, are thought to limit 
differentiation in favor of systems that maintain 
successful genotypes, the results obtained in this 
study suggest greater variability than is suspect- 
ed. Conclusions drawn from the performance of 
only one or a few genotypes out of a given 
population must be guarded, but it is possible 
that interactions of local populations with their 
environments are producing significant differ- 
entiation. Such differentiation would result ina 
flora better able to meet future exigencies than 
were it genetically uniform. 
Phytometer Study 
Procedures 
Three species were grown from seed in plas- 
tic pots, 20cm across and 21cm deep, filled 
with a mica peat mix and sunk to ground level at 
Palmer and Prudhoe in 1973. Seed of ‘‘Engmo” 
timothy, annual ryegrass, and ‘‘Sprite’’ peas 
were sown on the surface of the mix on June 26 
at both sites. The pots were fertilized about 
every week with 250 ml of Hoagland solution 
containing an additional 200 ppm of P. After 
germination the seedlings were reduced to five in 
each pot. The material was harvested 65 days 
from planting at Prudhoe and 66 days at Palmer. 
Various morphological and biomass measure- 
ments were made. 
