No. I.] SPERMATOGENESIS OF BATRACHOSEPS. 63 



to some extent at least, influences the formation of the leaders. 

 I refer to the fact that at the very beginning of the bouquet 

 stage those ends of the leaders which are not connected with 

 these chromoplasts point to the granosphere, and they not only 

 point to the sphere, but they actually cover a surface which is 

 just as wide as the granosphere, this fact determining the bou- 

 quet form. Just before the formation of the bouquet stage the 

 chromoplasts are situated in that part of the nucleus which is 

 nearest to the granosphere, but as the leaders are becoming 

 more defined, the chromoplasts move away from the spheres to 

 the opposite end of the nucleus, leaving the ends of the leaders 

 resting on that part of the nuclear wall nearest the spheres. 

 The ends of the leaders are so placed that at the point of con- 

 tact with the membrane they are closer together than a short 

 distance from it. If there thus exists a real influence on the 

 leaders from the spheres, this influence must be passing through 

 the nuclear membrane, which at this time, and for some time to 

 come, remains intact. The reduction in the number of chromo- 

 somes is certainly not performed by any rays or fibers, as these 

 have not yet penetrated the nuclear wall. 



The Radiosomic Process, or the Evolution of the Spheres, 

 Spindles, Fibers, ArcJiosome, and Accessory Archosomes. 



The radiosomic process begins in the polymorphous sper- 

 matogonia with the formation of the granosphere (Figs. 1-8). 

 The cytoplasm proper is then in the form of a very thin shell on 

 all sides surrounding the polymorphous nucleus. In this cyto- 

 plasmic envelop a denser area appears (Fig. 2), which is at first 

 homogeneous, but which later on differentiates into smaller 

 isolated areas or vacuoles, surrounded by denser staining 

 granules. At the same time an outer zone is forming of much 

 larger dimensions, but of less consistency. The outer zone 

 is the plasmosphere, and the inner one is the granosphere. 

 While this has taken place in the spheres, the cytoplasm has 

 spread over the larger part of the cell, having lost its thin 

 shell-like form. Already, with the first appearance of the 

 granosphere, there appears also in the cytoplasm one or more 



