EEPORT ON THE PHYLLOCARIDA. 37 



over, in all essential points rather w^ell with that found in Nebalia} The apparent 

 agreement with the Schizopoda in the mode in which the ova and embryos arc borne in 

 Nebalia during their development, is considerably lessened by the circumstance that 

 there is in the latter form no trace of any true incubatory pouch, the ova being simply 

 received within the valves of the carapace and kept in place by the aid of the branchial 

 legs. 



Concluding Remarls on the. Phylogeny of the Nebaliidx. — Owing to the suggestion 

 made by most naturalists, that Nebalia forms a direct transition between the Phyllopoda 

 and Podophthalmia, it has generally been supposed that the Nebaliidse have descended 

 from the Phyllopoda, and that, on the other hand, all the Podophthalmia should be 

 regarded as descendants from Nehalia-like ancestors. In his interesting treatise on the 

 phylogenetic relationship of the Malacostraca, Dr. Boas has sought to strengthen the 

 latter supposition by instituting a close comparison of the limbs in Nebalia with those 

 in the Malacostraca, and has thereby been led to the result, that the connecting link 

 between Nebalia and the great bulk of the Podophthalmia is represented by the 

 Euphausiidae, from which again all the other forms of that division are supposed to have 

 descended. It would seem that the chief reason that has led Dr. Boas to this view as 

 to the supposed close relationship between Nebalia and the Euphausiidse, is the apparent 

 agreement in the number of segments composing the anterior division of the body 

 (cephalothorax), and the uniformity in structure of the eight pairs of limbs succeeding 

 the oral parts in both. For in all other points the difference is in reality so very great, 

 as in most cases only to admit of the statement of a very general homology, such as 

 could also be made by comparing almost any forms of Crustacea. It has been stated 

 above that the resemblance of the branchial legs to the legs in the Euphausiidse is in the 

 genus Paranebalia considerably more pronounced than in Nebalia. But notwithstanding 

 this agreement, I still believe that there is no true relationship between the Nebaliidte 

 and the Euphausiidse, and that the above mentioned apparent conformity in the con- 

 struction of some of the limbs in both is merely accidental, a view that seems also to be 

 confirmed by the fact that in an otherwise closely related genus, Nebaliojjsis, these 

 limbs exhibit a form showing no similarity whatever to the corresponding limbs in the 

 Euphausiidse. If any true consanguinity had existed between the Nebaliida3 and the 

 Euphausiidae, the agreement between the two would certainly not be restricted to the 

 above named limbs, but might have been traced also in the rest of the organisation. But 

 this is by no means the case. On the contrary, it may easily be found on closer com- 

 parison, that the Euphausiidse in so far differ from the Nebaliidse even considerably 

 more than is the case with a great number of other Malacostraca. 



As to my own view on the probable phylogenetical relation of the Nebaliidge to other 



1 On Cyclestlieria liLslopi (Baird), a New Generic Type of Bivalved Phyllopoda, Forhandl. Vidensh Selsk. Christ., 

 1887 (to be shortly published). 



