THE MOUTHPARTS OF THE NEMATOCEROUS DIPTERA, II. 



BY VERNON L. KELLOGG, STANFORD UNIVERSITY, CALIFORNIA. 



In the following account of the 

 mouthparts of the Nematocera, that 

 nomenclature of parts is used which 

 represents the interpretation of the 

 mouthpart homologies most widely 

 accepted at the present time. As some 

 nomenclature is necessary and the 

 author's interpretation of the homol- 

 ogies may not, consistently with the 

 point of view adopted in this study, be 

 set out until the testimony of the study 

 has been presented, the nomenclature 

 of common acceptation is naturally 

 used. As the mouthparts of one repre- 

 sentative, at least, of each family, are 

 figured from drawings made by camera 

 lucida, the descriptions of the various 

 parts are made very brief. Owing to 

 the limitations of space, in most 

 instances the mouthparts are figured in 

 situ alone ; in some instances, however, 

 figures of two or three of the isolated 

 parts are given. As all the Nema- 

 tocerous mouthparts are alike in essen- 

 tial character and arrangement, the 

 figures of the cross sections of the 

 mouthparts of Blepharocera capitata 

 illustrate nearly as well the conditions 



presented by any other of the mandible- 

 possessing Nematocerous females. 



Blepijaroceridae. 



Liponeura ? sp. This species is a 

 large undescribed Blepharocerid, taken 

 by me in California, and should proba- 

 bly be made the type of a new genus. 

 The mouthparts of the female consist of 

 the following well developed, independ- 

 ent and easily distinguishable parts 

 shown in figure i in situ to reveal the 

 relative size and natural position with 

 regard to each other : a labrum-epi- 

 pharynx (/. ep), a pair of mandibles 

 {nut), a pair of maxillae (?«.v), a labium 

 (//') and a hypopharynx {/np) . 



The labrum-epipharynx (fig. i A, 1. 

 ep) is elongate, slender, and pointed, 

 and bears numerous taste pits (?) on its 

 lower (inner) surface which is concave. 



The mandibles (fig. i A, nui) are long, 

 slender, well chitinized, and each is 

 sharply, finely and conspicuously serrate 

 along the distal half of the inner mar- 

 gin. The mandibles are articulated 

 widi the head capsule wholly distinctly 

 from the other mouthparts. 



