852 QUADRULA 
Unio keinerianus Lea, Pr. Am. Phil. Soc., V, 1852, p. 251; 
Tr: Am. Phil; Soc. X; 1852, p.281, plax xt, se. 4aOwelps. 
V, 1852, p. 37, pl. xx111, fig. 40. 
Margaron (Unio) kemerianus L&A, Syn., 1852, p. 22. 
Margaron (Unio) kienerianus Lea, Syn., 1870, p. 34. 
Quadrula pustulosa var. keimeriana SIMPSON, Syn., 1900, p. 
780. 
Seems tc merge into variety pernodosa, but it is smaller 
and the epidermis is differently colored. Some specimens have 
broken green bands and are feebly rayed. 
This form was named by Lea after Kiener, but he inad- 
vertently misspelled the name. It is therefore proper to cor- 
Tech tne eLLor. 
QUADRULA COOPERIANA (Lea). 
Shell subtriangular, rarely suborbicular, subinflated, solid, 
inequilateral; beaks very high, rather full, turned forward 
over a deep lunule; posterior ridge low, rounded; anterior 
end usually obliquely truncated above, rounded below; base 
line more or less rounded; outline of dorsal slope slightly 
curved, sometimes raised almost into an angle behind the 
ligament; surface with irregular growth lines; posterior two- 
thirds covered with strong, irregular pustules; epidermis red- 
dish-brown; pseudocardinals triangular, not greatly rough- 
ened ; lateral double in the right valve; beak cavities deep, com- 
pressed; muscle scars deep; nacre white or pink, thinner be- 
hind where it is often bronzy. 
Length 90, height 80, diam. 47 mm. 
Length &o, height 84, diam. 43 mm. 
Ohio, Cumberland, and Tennessee River systems. Reported 
by Keyes at Muscatine, lowa, but this is probably an error. 
Type locality, Ohio river. 
Unio cooperianus Lea, Tr. Am. Phil. Soc., V, 1834, p. 61, pl. 
vill, fig. 21; Obs., I, 1834, p. 173, pl. vit, fig. 21—HANLEY, 
Biv. Shells, 1843, p. 180, pl. xxi, fig. 1.—Kuster, Conch. 
Cab. Unio, 1861, p. 183, pl. Lv, fig. 5. 
Margarita (Unio) cooperianus Lea, Syn., 1836, p. 16; 1838, 
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