REPORT ON THE CRINOIDEA. 215 
With regard to the support beneath the radial pentagon of Hudesicrinus, I see no 
reason for doubting that the upper part consists of united basals. In fact, one of de 
Loriol’s figures! shows a distinct horizontal sutural line crossing the middle of the support, 
and separating the infra-radial portion with interradial crests on its upper surface from 
the more spreading, attached part below. In another specimen this suture seems to be 
indicated by an external circular ridge ; but the upper face of the support is marked by 
five petaloid depressions, one of them considerably larger than the rest, which surround 
acentral pit. De Loriol, and, I think, rightly so, regards these depressions as corresponding 
to the cavities of the chambered organ, the largest being that of the large radial in the 
trivium. Similar but more recular depressions appear round the middle of the upper 
surface of the large basal pentagon in Apiocrinus milleri, Quenstedt ; and they are also 
shown in de Loriol’s figure of Apiocrinus roissyanus, while it will be remembered that 
the chambered organ is invariably in close relation with the basals (Pl. VITb. figs. 1, 2; 
Pl; XXIV. figs. 6, 7; Pl. LVIII. figs. 1, 3—ch; Pl. LXI.). There can then, I think, be 
no doubt as to the presence of basals in Hudesicrinus, so that the so-called support does 
not in reality differ essentially from the centro-dorsal of Cotylecrinus. The radials, how- 
ever, are very different in the two cases.. Those of Cotylecrinus are equal and similar ; 
but in Eudesicrinus they are thus described by de Loriol,’ “Ces pieces sont fort inégales; 
lY'une est notablement plus longue et plus large que les autres, convexe et un peu arquée 
en dehors, mais d’une maniére uniforme ; les deux qui la touchent, bien plus courtes et plus 
étroites qu’elle-méme, ne sont guére plus larges, mais plus longues que les deux autres ; 
ces derniéres, qui sont les plus courtes et placées vis-A-vis de la plus large, s’arquent en dedans 
et se coudent un peu vers leur bord supérieur. Dans les échantillons frais la surface externe 
est couverte de granules épars, écartés, plus ou moins gros et plus ou moins serrés.” 
Here then we have a type which bears an unusually close resemblance to Holopus. 
The calyx is higher on one side than on the other owing to the inequality of the radials, 
the central one of the trivium being the largest, while the outer surface, not only of the 
radials, but also of the arm-joints, is coarsely granular or tubercular. Eudesicrinus, however, 
differs from Holopus in one or two minor points. The fossee on the radials which lodged 
the dorsal ligaments are larger; and there seems to have been a true muscular joint be- 
tween the second and the axillary radials, a character presented by no recent Crinoid, and 
also, so far as I know, by no other fossil species. In Hugeniacrinus mayalis from the 
same horizon the two joints are united by syzygy, just as I believe to be the case in 
Holopus, while the calyx is less.coarsely granular. These two species are quite small 
relatively to Holopus, the radials of Hudesicrinus not reaching a height of more than 
11mm. Associated with them in the Leptena-bed (Middle to Upper Lias) of Calvados, in 
Normandy, are some wedge-shaped arm-joints with the muscle-plates on their broad outer 
sides produced into strong upward processes. These joints have very much the appear- 
1 Paléont. Frang., loc. cit., pl. 29, fig. 7.  * Ibid., pl. 44, fig. 2, 6. 3 [bid., p. 78. 4 Ibid., pl. 11, figs. 1-4. 
