30 
also pointed out to me the strong similarity of the males of Dorylus 
to those of the wasps of the East Indian Vespa doryloides Saussure." 
The scant evidence thus far obtained seems to show that a colony 
of drivers has only one large, distended queen, like that of the Ter- 
mites. Emery notes that the last joints of the legs are usually 
broken off in the specimens known to. him.’ Such mutilation may 
well arise while the huge creature is being dragged about, as stated 
by the native Africans. 
The interesting feature of the drivers, from the standpoint of the 
study of the kelep, lies in the fact that the differences between them 
and the true ants are in several respects the same, though much more 
accentuated than between the ants and the kelep. Thus, interbreeding 
is accomplished in both groups by the circulation of the males, the 
females, apparently, never emerging from the nests. The drivers 
carry their larve, and presumably their queens also, for long dis- 
tances, their close ranked, rapidly moving columns sometimes requir- 
ing several hours to pass a given point. In food habits, too, they 
are carnivorous, like the keleps. They not only capture insects and 
other arthropods, but young or helpless vertebrates also. Active 
creatures even, like rats and mice, are often secured, the pain inflicted 
by the bites of the drivers causing them to roll about on the ground 
and thus permit more and more of their diminutive enemies to 
attack them. Any animal too large to be carried away by a single 
worker is cut in pieces. The kelep follows the more social plan of 
moving large insects and centipedes by the combined strength of 
several workers. The drivers are also like keleps in having a sting 
which they do not use as a weapon of offense against man, but they 
bite viciously and will not let go, even though their bodies be pulled 
loose from their heads. 
The association of the keleps with the drivers permits a better 
appreciation of their habits in two respects. First, the apparently 
high development of their instinct of place or ability to find their 
way about, even in new situations. The drivers are fierce warriors, 
a Doctor Ashmead informs me that the males of the tribe Amblyponine, near 
relatives of the kelep, are very closely similar to those of the Mutillidz, even to 
the structure of the petiole and the venation of the anterior wings. 
The flying to lights of the males of the Dorylide, or drivers, is also shared by 
the males of Myrmosa and related genera of parasitic wasps formerly placed in 
the-Mutillidze, but which Doctor Ashmead now separates as a distinct family, 
Myrmoside, intermediate between the Mutillidz and Vespidz, but having wing- 
less females like the Mutillidze and the Dorylids. Some of the Bethylidz also 
have wingless females, but these are not supposed to have any close affinities 
with the other families. The Thynnidze, which have been thought to connect 
with the Mutillidse and the ants, likewise have wingless females. Finally, a 
few of the Mutillidze are wingless in both sexes. 
b Emery, E., 1895, Ann. Soc. Entom., France, 64:75. 
