UPPER TENNESSEE DRAINAGE. 541 
Rock River, Flint River in Madison Co., Elk River, Shoals Creek, 
Bear Creek). 
15. QUADRULA METANERVA (Rafinesque), 1820. 
Obliquaria metanerva Rafinesque, ’20.—Unio metanerva Lewis, ’71. 
—Quadrula metanerva Ortmann, 712), p. 255 (anatomy) .— 
Quadrula metanerva Simpson, ’14, p. 834. 
In the Tennessee at and below Knoxville. Above Knoxville, I 
found only a single specimen in the Holston, at Mascot, Knox Co. 
Rare. 
Type locality: Kentucky River (according to Vanatta, ’15, the 
types are from Ohio River). 
16. QUADRULA INTERMEDIA (Conrad), 1836. 
Unio intermedius Conrad, ’36.—Unio tuberosus Lea, ’40.—Unio 
sparsus Lea, ’41—Unio intermedius, sparsus, tuberosus Lewis, 
’71.—Quadrula sparsa Ortmann, ’12b (anatomy ).—Quadrula in- 
termedia Goodrich, *13, p. 93——Quadrula tuberosa, tuberosa 
Sparsa, intermedia Simpson, ’14, pp. 836, 837. 
According to Simpson, Q. tuberosa is more swollen than Q. in- 
termedia, and I have seen rather swollen specimens in the Walker 
collection from the Cumberland River and the Tennessee in North 
Alabama. However, the original figure of U. tuberosus Lea is not 
much swollen. My specimens from the upper Tennessee drainage 
are all more or less compressed. We might have here again a case 
where in larger rivers a more swollen form turns up. But even if 
this should be correct, we should only regard these forms as races 
of the same species. 
Reported from Nolichucky, Holston, Clinch, and Tennessee 
rivers, and said to be abundant. According to the material collected 
and seen by myself, it is decidedly a rare species. I know it from the 
Holston at Church Hill, Hawkins Co., Tenn.; from:the South Fork 
Holston at Pactolus and Bluff City, Sullivan Co., Tenn.; from North 
Fork Holston at Mendota, Washington Co., Va.; and from the 
Clinch River at Clinchport, Scott Co., Va. (Walker coll.), and 
Cleveland, Russell Co., Va. 
