Occasional Papers of the Musewm of Zoology 17 
QUADRULA NODULATA Rafinesque, 1820 
Type locality: Kentucky River. 
Obliquaria (Quadrula) nodulata, Rafinesque, ’20, p. 307, pl. 81, f. 
17-18. 
Unio pustulatus Lea, “31, p. 79, pl. 7, f. 9: 
U. nodulatus Raf. = U. pustulatus Lea, Conrad, ’34, p. 70; ’37, p. 80. 
Quadrula pustulata (Lea), Simpson, ’14, p. 856. 
Quadrula nodulata (Raf.) =Q. pustulata (Lea), Vanatta, ’15, p. 
557 (“type” examined). 
Quadrula nodulata (Raf.), Utterback, ’16, p. 57. 
Obli. nodulata Raf. =Q. pustulata (Lea), Walker, ’18¢, p. 168 (“if 
identifiable”). 
The original description of nodulata mentions and the 
(poor) figure shows the characteristic posterior “wing” of 
this shell; also the arrangement of the distant (text) tubercles 
in two rows is suggested in the figure. A nodulous shell with 
these characters can only be the pustulata of Lea. Thus, 
Rafinesque’s species is recognizable, which is confirmed by 
Vanatta’s examination of the so-called Rafinesque-Poulson 
~ types: 
QUADRULA METANEVRA Rafinesque, 1820 
Type locality: Kentucky River. 
Obliquaria (Quadrula) metanevra Rafinesque, ’20, p. 305, pl. 81, 
f. 15-16. 
Unio nodosus Barnes, ’23, p. 124, pl. 6. 
U. metanevra Raf. =U. nodosus Bar., Conrad, ’34, p. 70; Ferussac, 
a5seDs 27.9 Conrady 2s5=p. lOs plate te 2 
Theliderma metanevra (Raf.), Swainson, ’40, p. 378. 
Quadrula metanevra (Raf.), Agassiz, ’52, p. 48; Simpson, 14, p. 834. 
There has never been any doubt, since the time of Conrad, 
as to the identity of this species, and indeed Rafinesque’s 
description and figure are identifiable. 
However, this species should not be regarded as the type of 
the genus Quadrula. It belongs in the group of U. cylindricus 
Say, for which Orthonymus Agassiz ('52) is to be used as 
sectional (or subgeneric) name (see above). 
