HAZEN: NEW SPECIES OF LOBOMONAS 135 
ferred to. This theory that amoeboid movemnt is due to alter- 
ations of the colloidal state is only in the nature of confirm- 
ation and extension of the view advanced more than forty 
years ago by Montgomery (17, 18) that protoplasmic movement 
of amoeboid organisms consists in ‘an alternate expansion and 
contraction of organic substance’: Montgomery even anticipated 
Hyman in expressing the idea that the liquefaction which occas- 
ions pseudopodium advancement is itself due to metabolic changes, 
while he anticipated Rhumbler (19) in the idea of a non-homo- 
geneity of the primitive protoplasmic mass which permits various 
functions to be carried on in different regions at the same 
time. 
This somewhat lengthy excursus (which is yet only the 
briefest possible summary of a voluminous literature) has been 
introduced here only to bring before botanists, to whom the field 
may be unfamiliar, facts which it is believed may be directly 
applied in the case of the chlamydomonads we are considering. 
The form development of Lobomonas, Brachiomonas, and Ptero- 
monas must be essentially amoeboid for-a brief period during 
the organization of the daughter cells, and we are justified 
in assuming that their lobes and excrescences are the expression 
of the same non-homogeneous organization of the protoplast 
as is characteristic of Amoeba. 
For this view, furthermore, we may draw an additional 
parallel from the results of microdissection. In the developing 
oogonium of Fucus, Seifriz reports (21) that the viscosity of the 
protoplasm changes from liquid consistency in the young uni- 
nucleate stage to slightly viscous consistency when the division 
into eight eggs is just complete, and to decidedly viscous consis- 
tency (just under the viscosity of glycerine) in the mature nor- 
mally discharged egg: that is, in Seifriz’s scale of ten grades of 
viscosity—the first attempt on the part of microdissectionists 
at standardization in this matter—the variation is from grade 
3 to grade 6. Yet further, from the behavior of disintegrating 
eggs of Fucus, Seifriz (20) finds that the process may be localized 
in such a manner as to indicate ‘a gross structure of the egg 
plasm, i. ¢., the protoplasm is composed of many centers of activity 
in which different chemical reactions take place.’ [have recently 
found a condition almost precisely similar in the case of a newly 
discovered polyblepharid genus which it is hoped may soon be 
published. The cells of this species, though surrounded only 
by an exceedingly delicate protoplasmic membrane, are never- 
